Niphotrichum canescens

(Hedwig) Bednarek-Ochyra & Ochyra in R. Ochyra et al.

in R. Ochyra et al., Cens. Cat. Polish Mosses, 137. 2003,.

Basionym: Trichostomum canescens Hedwig Sp. Musc. Frond., 111. 1801
Synonyms: Bryum canescens (Hedwig) Withering Grimmia canescens (Hedwig) Müller Hal. Racomitrium canescens (Hedwig) Bridel
Treatment appears in FNA Volume 27. Treatment on page 287. Mentioned on page 286, 288, 292.

Plants medium-sized and fairly slender to large, in loose or dense patches or tufts, dull green, yellow-green or yellowbrown, often hoary when dry. Stems (1–) 3–10 (–12) cm, sparingly or profusely irregularly branched or often regularly pinnately branched with short, tuft-like branchlets. Leaves straight to distinctly falcate, imbricate when dry, patent to recurved when moist, ovate to broadly ovatelanceolate, (1.5–) 2–2.8 (–3.3) × 0.8–1.3 mm; margins broadly recurved to revolute throughout; apices rather abruptly narrowed, hyaline or not, sometimes muticous, usually awned, awns broad and stout to subulate, not reflexed, not or distinctly decurrent, mostly serrulate and spinulose, papillose distally and strongly so basally; costa weak, extending 1/2–3/4 way up the leaf, flattened and weakly convex abaxially, 90–120 µm wide near the base; basal-cells rectangular, 30–50 × 4–7 µm strongly papillose with large papillae, often epapillose in 1–3 rows at the insertion; alar cells usually rounded and inflated in 3–5 rows, thin-walled, hyaline to yellowish hyaline, forming prominent, decurrent, somewhat inflated auricles; supra-alar cells elongate, thin and straight-walled, producing a pellucid marginal border of 20–30 cells; medial and distal laminal cells rectangular, 10–30 × 5–10 µm, strongly papillose. Inner perichaetial leaves membranous, hyaline, acute, usually with hyaline awns. Seta dark red to reddish-brown, 5–25 mm. Capsule brown, narrowly ellipsoid to cylindric, 1.5–2.5 mm, smooth to somewhat furrowed when dry; peristome teeth 600–800 µm, reddish-brown, densely papillose. Spores 8–11 µm.

Discussion

Subspecies 2 (2 in the flora).

Niphotrichum canescens is separated from all other congeners by its leaves broadly canaliculate to obtusely keeled in the distal portion, stout awns that are strongly papillose throughout and weakly serrulate, and short and forked costae ending well before the apex, usually in mid leaf or at three quarters of the leaf length.

Selected References

None.

Key

1 Plants mostly robust; leaves ovate to broadly ovate-lanceolate, hyaline at the apex, strongly obtusely keeled and cucullate distally; awns non-decurrent. Niphotrichum canescens subsp. canescens
1 Plants fairly gracile; leaves ovate-lanceolate to broadly ovate-lanceolate, chlorophyllose at the apex, less obtusely keeled in the upper part; awns decurrent. Niphotrichum canescens subsp. latifolium
... more about "Niphotrichum canescens"
not +  and hyaline +
inflated +  and decurrent +
Ryszard Ochyra +  and Halina Bednarek-Ochyra +
(Hedwig) Bednarek-Ochyra & Ochyra in R. Ochyra et al. +
capillaceous +
spinulose +  and serrulate +
flexuose +  and straight +
not reflexed +
papillose +
decurrent +  and subulate +
nodulose;oblate;quadrate;elongate +
rectangular +
divided +
Trichostomum canescens +
2;3;2;numerous +
papillose +
linear-subulate +  and tuft-like +
undifferentiated +
naked +, plicate +  and smooth +
lobed +, lacerated +, mitrate-campanulate +, mitrate +  and cucullate +
small;large +
furrowed +  and smooth to somewhat +
narrowly ellipsoid;cylindric +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
straight-walled +  and thin-walled +
hyaline +  and yellowish hyaline +
20 (?) +  and 30 (?) +
elongate +, inflated +  and rounded +
enlarged +
subpercurrent +
1/2 +  and 3/4 +
convex +  and flattened +
90um +  and 120um +
North America +  and Eurasia +
0.28 cm2.8 mm <br />0.0028 m <br /> (0.33 cm3.3 mm <br />0.0033 m <br />) +
thick-walled +  and sinuose +
straight +
2mm;2.8mm +
ovate +  and broadly ovatelanceolate +
0.08 cm0.8 mm <br />8.0e-4 m <br /> (0.13 cm1.3 mm <br />0.0013 m <br />) +
muticous +, acute +  and rounded-obtuse +
long-subulate +
piliferous;obtuse;acute +
cribrose +  and perforated +
lanceolate;linear +
in R. Ochyra et al., Cens. Cat. Polish Mosses, +
dark red +  and reddish-brown +
sinistrorse +  and erect +
papillose +  and smooth +
short to long +
0.5 cm5 mm <br />0.005 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
ornamented +  and smooth +
globose +
3 cm30 mm <br />0.03 m <br /> (10 cm100 mm <br />0.1 m <br />) +
Bryum canescens +, Grimmia canescens +  and Racomitrium canescens +
Niphotrichum canescens +
Niphotrichum sect. Niphotrichum +
species +
divided +
yellowbrown;yellow-green;green;dull +
fairly slender;large +