Pellaea glabella subsp. simplex

(Butters) A. Löve & D. Löve

Taxon 26: 325. 1977.

IllustratedEndemic
Basionym: Pellaea glabella var. simplex Butters Amer. Fern J. 7: 84. 1917
Synonyms: Pellaea atropurpurea var. simplex (Butters) C. V. Morton Pellaea occidentalis subsp. simplex (Butters) Gastony Pellaea suksdorfiana
Treatment appears in FNA Volume 2.

Ultimate segments essentially glabrous; sporangia containing 32 spores; spores averaging 60–72 µm diam. n = 2n = 116, apogamous.


Phenology: Sporulating summer–fall.
Habitat: Calcareous cliffs and ledges, usually on limestone
Elevation: 900–3000 m

Distribution

V2 66-distribution-map.gif

Alta., B.C., Ariz., Colo., Idaho, Mont., N.Mex., Utah, Wash., Wyo.

Discussion

This western counterpart of Pellaea glabella subsp. glabella is an apogamous tetraploid. A. F. Tryon (1957) and D. B. Lellinger (1985) hypothesized that it might have arisen as a hybrid between the western diploid member of the P. glabella complex (here called subsp. occidentalis) and P. atropurpurea. G. J. Gastony (1988) has shown conclusively, however, that P. glabella subsp. simplex is an autopolyploid derivative of subsp. occidentalis and does not contain genes contributed by P. atropurpurea.

Selected References

None.

Lower Taxa

None.
Michael D. Windham +
(Butters) A. Löve & D. Löve +
cordate +, truncate +  and rounded +
Pellaea glabella var. simplex +
glabrous +  and pubescent +
linear-oblong +  and ovatelanceolate +
1cm;8cm +
erose-denticulate +
straight +
Alta. +, B.C. +, Ariz. +, Colo. +, Idaho +, Mont. +, N.Mex. +, Utah +, Wash. +  and Wyo. +
900–3000 m +
yellow +  and white +
obcordate;reniform +
Calcareous cliffs and ledges, usually on limestone +
monomorphic +
clustered;widely scattered +
recurved +  and reflexed +
noncircinate +  and circinate +
2 cm20 mm <br />0.02 m <br /> (40 cm400 mm <br />0.4 m <br />) +
? (?) +  and ? (?) +
entire +, erose +  and dentate +
entire +  and denticulate +
pubescent +  and glabrous +
Sporulating summer–fall. +
reddish-brown +
persistent +
occasional +
hairlike +  and linear-subulate +
thin +, 0.1mm +  and 0.3mm +
Illustrated +  and Endemic +
brown +  and tan +
cristate +  and tetrahedral-globose +
dark colored +
0.5 cm5 mm <br />0.005 m <br /> (1 cm10 mm <br />0.01 m <br />) +
Pellaea atropurpurea var. simplex +, Pellaea occidentalis subsp. simplex +  and Pellaea suksdorfiana +
Pellaea glabella subsp. simplex +
Pellaea glabella +
subspecies +
0.5 cm5 mm <br />0.005 m <br /> (2 cm20 mm <br />0.02 m <br />) +
leathery +  and herbaceous +
crescent--shaped +  and roundish +
branched +  and anastomosing +