Pennellia

Nieuwland

Amer. Midl. Naturalist 5: 224. 1918.

Etymology: For Francis Whittier Pennell, 1886–1952, American botanist
Basionym: Heterothrix (B. L. Robinson) Rydberg Bull. Torrey Bot. Club 34: 435. 1905,
Synonyms: Lamprophragma O. E. Schulz
Treatment appears in FNA Volume 7. Treatment on page 559. Mentioned on page 224, 226, 235, 243, 257, 561.
 TaxonIllustrator 
FNA7 P69 Pennellia tricornuta.jpegPennellia tricornuta
Hesperis matronalis
Iberis sempervirens
Yevonn Wilson-Ramsey
Yevonn Wilson-Ramsey
Yevonn Wilson-Ramsey

Perennials; not scapose; pubescent or glabrous, trichomes simple, with stalked, forked or dendritic ones. Stems erect [ascending], often branched distally. Leaves basal and cauline; petiolate or sessile; basal (withered with age), rosulate, petiolate, blade margins entire, dentate to sinuate, or runcinate; cauline subsessile or sessile [shortly petiolate], blade margins entire [dentate to sinuate]. Racemes (several-flowered, sometimes secund, lax), considerably elongated in fruit. Fruiting pedicels erect, descending, ascending, or divaricate-ascending, slender, (glabrous). Flowers (usually actinomorphic, rarely zygomorphic, cupshaped); sepals erect, oblong to ovate, lateral pair slightly to strongly saccate basally, (usually pubescent, rarely glabrous); petals white (lavender or purple apically), spatulate to oblanceolate, (± longer than sepals), claw undifferentiated from blade, (apex obtuse); stamens slightly tetradynamous; filaments dilated basally; anthers ovate to oblong, (apex obtuse); nectar glands confluent, subtending bases of stamens. Fruits siliques, sessile or shortly stipitate, linear, not torulose, straight or curved, terete or latiseptate; valves each with prominent or obscure midvein, glabrous or pubescent; replum rounded; septum complete; ovules 40–250 per ovary; style distinct or obsolete; stigma capitate. Seeds uniseriate or biseriate, plump or flattened, winged or not, oblong to ovate; seed-coat (reticulate-areolate), mucilaginous or not when wetted; cotyledons accumbent or incumbent. x = 8.

Distribution

sw United States, Mexico, Central America, South America

Discussion

Species 8 (3 in the flora).

Key

1 Fruits strongly latiseptate, 2-2.5 mm wide; seeds 2.4-2.8 mm, winged (broadly); septums opaque; ovules 40-50 per ovary; cotyledons accumbent. Pennellia tricornuta
1 Fruits terete or slightly latiseptate, 0.6-1.8 mm wide; seeds 0.7-1.3 mm, not winged; septums transparent; ovules 90-250 per ovary; cotyledons incumbent > 2
2 Racemes not secund; fruiting pedicels straight, erect or ascending; fruits (1.5-)2.2-5.8 cm, terete; ovules 90-140 per ovary; sepals 2.7-4 mm; petals 1.5-4(-5) mm. Pennellia micrantha
2 Racemes secund; fruiting pedicels arcuate, reflexed; fruits 5-10 cm, latiseptate; ovules 150-250 per ovary; sepals 4-9 mm; petals (4-)5-7(-12) mm. Pennellia longifolia

"elongated" is not a number."thick" is not a number.

... more about "Pennellia"
ovate +  and oblong +
Sara Fuentes-Soriano +  and Ihsan A. Al-Shehbaz +
Nieuwland +
Heterothrix +
appendaged +  and unappendaged +
not auriculate +
dentate +  and sinuate or runcinate +
much smaller +  and reduced +
urceolate +, campanulate +  and tubular +
branched +  and simple +
sessile +  and subsessile +
well-developed +
undifferentiated +
not +  and differentiated +
emarginate +  and entire +
incumbent +  and accumbent +
sw United States +, Mexico +, Central America +  and South America +
straight +  and curved +
For Francis Whittier Pennell, 1886–1952, American botanist +
not winged +  and unappendaged +
connate +  and distinct +
zygomorphic +  and actinomorphic +
2-valved +  and capsular +
latiseptate +  and unsegmented +
angustiseptate +  and terete +
latiseptate +, stipitate +  and sessile +
curved +  and straight +
terete +  and not torulose +
sessile +  and petiolate +
alternate +, not +  and rosulate +
cauline +  and basal +
connate +  and distinct +
decurrent +
lobed;dentate;lobed;dentate +
obscure +  and prominent +
tenuinucellate +, crassinucellate +  and bitegmic +
campylotropous +  and anatropous +
rudimentary +
spatulate +  and oblanceolate +
trinucleate +  and 3(-11)-colpate +
Amer. Midl. Naturalist +
divaricate-ascending +, ascending +, descending +  and erect +
slender +
fuentes-soriano2004a +  and rollins1980a +
not +  and mucilaginous +
not +  and winged +
biseriate +  and uniseriate +
not +  and mucilaginous +
white +, black +, brown +  and yellow +
oblong +  and ovate +
persistent +  and caducous +
distinct +
oblong +  and ovate +
reduced +
latiseptate +, schizocarpic +, samaroid +  and lomentaceous +
indehiscent +  and dehiscent +
angustiseptate +, flat +, angled +, terete +, torulose +, not +, segmented +  and nutletlike +
tetradynamous +
procumbent +  and decumbent +
2-lobed +, entire +, conical +  and capitate +
subsessile +, sessile +  and petiolate +
persistent +
distinct +
Lamprophragma +
Pennellia +
Brassicaceae tribe Halimolobeae +
coiled +, 1-7-veined +  and veined +
pubescent +  and glabrous +
anastomosing +
glabrous +  and pubescent +
rhizomatous +, taprooted +, scapose +  and not +
perennial +, biennial +  and annual +
subshrub +  and herb +
aquatics +  and terrestrial +
glabrous +  and pubescent +