Penstemon procerus

Douglas ex Graham

Edinburgh New Philos. J. 7: 348. 1829.

Common names: Small-flower beardtongue
IllustratedEndemic
Synonyms: Penstemon confertus var. procerus (Douglas ex Graham) Coville
Treatment appears in FNA Volume 17. Treatment on page 216. Mentioned on page 184, 186, 187, 213, 220.

Herbs. Stems ascending to erect, 3–55 (–70) cm, glabrous or ± retrorsely hairy, not glaucous. Leaves basal and cauline, basal sometimes poorly developed, ± leathery or not, glabrous, puberulent, or retrorsely hairy; basal and proximal cauline 9–90 (–115) × 2–19 mm, blade spatulate to oblanceolate, lanceolate, or elliptic, base tapered, margins entire, apex rounded to obtuse or acute; cauline 2–6 pairs, sessile or proximals sometimes short-petiolate, 6–85 × 1–17 mm, blade oblanceolate to spatulate, obovate, oblong, elliptic, lanceolate, or linear, base tapered to clasping, margins entire, apex rounded to obtuse or acute. Thyrses interrupted, cylindric, 0.5–23 cm, axis glabrous or retrorsely hairy, verticillasters 1–8 (–11), cymes 3–11-flowered, 2 per node; proximal bracts lanceolate to linear, 10–50 (–60) × 1–8 (–16) mm, margins entire; peduncles and pedicels erect, glabrous, puberulent, or retrorsely hairy. Flowers: calyx lobes ovate to obovate or lanceolate, 1.2–5 (–5.5) × 0.5–2 mm, apex truncate, obtuse, acute, acuminate, cuspidate, caudate, or mucronate, glabrous or puberulent to retrorsely hairy; corolla violet to blue or purple, with, sometimes without, reddish purple nectar guides, funnelform, 7–11 mm, glabrous externally, moderately white or yellow-lanate internally abaxially, tube 2–3 mm, throat slightly inflated, 2–3 mm diam., 2-ridged abaxially; stamens included, pollen-sacs opposite, explanate, 0.3–0.6 mm, dehiscing completely, connective splitting, sides glabrous, sutures smooth or papillate; staminode 4–6 mm, included, 0.2–0.3 mm diam., tip straight to slightly recurved, distal 0.5–1 mm pilose, hairs yellow or golden yellow, to 0.6 mm, rarely glabrous; style 4–7 mm. Capsules 3–5 × 2–4 mm, glabrous.

Distribution

Alta., B.C., Man., Sask., Yukon, Alaska, Calif., Colo., Idaho, Mont., N.Dak., Nev., Oreg., Utah, Wash., Wyo.

Discussion

Varieties 6 (6 in the flora).

Penstemon procerus is a widespread, frequent, and highly variable species distributed throughout much of western North America. This variability has resulted in varied interpretations of its taxonomic units.

Selected References

None.

Key

1 Calyx lobes (2–)3–5(–5.5) mm, apices acuminate, cuspidate, or short- to long-caudate. > 2
2 Verticillasters (1–)3–8(–11); stems (6–)15–55(–70) cm; thyrses (1–)5–23 cm. Penstemon procerus var. procerus
2 Verticillasters 1 or 2(–5); stems 3–20(–28) cm; thyrses 1–5(–13) cm. Penstemon procerus var. tolmiei
1 Calyx lobes 1.2–3.2 mm, apices truncate to obtuse, acute, acuminate, mucronate, or short-caudate. > 3
3 Verticillasters 1 or 2(–5); stems 3–15(–20) cm; cauline leaves 1–4(–8) mm wide. Penstemon procerus var. formosus
3 Verticillasters (1 or)2–6; stems (4–)8–35(–42) cm; cauline leaves 2–13 mm wide. > 4
4 Calyx lobes 1.2–2.4 mm, apices truncate to obtuse or mucronate, rarely short-caudate or acute; Nevada. Penstemon procerus var. modestus
4 Calyx lobes (1.4–)1.8–3.2 mm, apices acute to acuminate or short-caudate; California, Oregon, Utah. > 5
5 Pollen sacs 0.3–0.4 mm; corollas 7–10 mm; stems 10–35(–42) cm; Utah. Penstemon procerus var. aberrans
5 Pollen sacs 0.4–0.6 mm; corollas 9–12 mm; stems 9–20(–30) cm; California, Oregon. Penstemon procerus var. brachyanthus

"/4+timescorollathroat" is not declared as a valid unit of measurement for this property.

... more about "Penstemon procerus"
puberulent +  and retrorsely hairy +
mucronate +, caudate +, cuspidate +, acuminate +, acute +, obtuse +, truncate +, rounded +  and obtuse or acute +
Craig C. Freeman +
Douglas ex Graham +
hairy +  and glabrous +
alternate +, subopposite +, whorled +  and opposite +
glaucescent +
alternate +, opposite +, whorled +, helical +  and subopposite +
cauline +, proximal +  and basal +
rarely +, not glaucous +, glabrate +, pubescent +, hairy +, glandular-lanate +, glandular-pubescent +, puberulent +, glandular +  and glabrous +
hairy +, puberulent +  and glabrous +
not +  and leathery +
not +  and leathery +
tapered;tapered +
oblanceolate;spatulate obovate oblong elliptic lanceolate or linear +
not +  and leathery +
0.5 cm5 mm <br />0.005 m <br /> (0.55 cm5.5 mm <br />0.0055 m <br />) +
ovate;obovate or lanceolate +
short-tubular +  and short-campanulate +
0.3 cm3 mm <br />0.003 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
petiolate +  and sessile +
Small-flower beardtongue +
personate +, not personate +, bilabiate +  and symmetric +
violet +  and blue +
glandular-pubescent +, hairy +, glabrous +, , +, glabrescent +  and glabrate +
ventricose +, tubular +  and tubular-funnelform +
ampliate +
3-11-flowered +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
scale-like +  and not needlelike +
Alta. +, B.C. +, Man. +, Sask. +, Yukon +, Alaska +, Calif. +, Colo. +, Idaho +, Mont. +, N.Dak. +, Nev. +, Oreg. +, Utah +, Wash. +  and Wyo. +
drupe-like +
curved +  and straight +
1 +  and 4 +
golden yellow +  and yellow +
0 cm0 mm <br />0 m <br /> (0.06 cm0.6 mm <br />6.0e-4 m <br />) +
membranous +
terminal +  and axillary +
persistent +  and deciduous +
cauline +  and basal +
orbiculate;linear;orbiculate;linear;truncate;oblanceolate;obovate;oblong;elliptic;round;lanceolate;ovate +
glutinous +
glandular-ciliolate +, hairy +, puberulent +, glabrescent +, glabrous +  and glandular-pubescent +
lacerate +, toothed +  and entire +
white +  and reddish purple +
yellow-lanate +  and glabrous +
0.7 cm7 mm <br />0.007 m <br /> (1.1 cm11 mm <br />0.011 m <br />) +
axile +  and parietal +
basal +, apical +  and superior +
tenuinucellate +, unitegmic +  and hemitropous +
campylotropous +, hemianatropous +  and anatropous +
hairy +, puberulent +  and glabrous +
hairy +, puberulent +  and glabrous +
5 +  and 4 +
navicular-sigmoid +
0.03 cm0.3 mm <br />3.0e-4 m <br /> (0.06 cm0.6 mm <br />6.0e-4 m <br />) +
1/5 +  and 2/3 +
short-petiolate +
5 cm50 mm <br />0.05 m <br /> (6 cm60 mm <br />0.06 m <br />) +
0.8 cm8 mm <br />0.008 m <br /> (1.6 cm16 mm <br />0.016 m <br />) +
1 cm10 mm <br />0.01 m <br /> (5 cm50 mm <br />0.05 m <br />) +
lanceolate +  and linear +
0.1 cm1 mm <br />0.001 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
0.6 cm6 mm <br />0.006 m <br /> (8.5 cm85 mm <br />0.085 m <br />) +
0.1 cm1 mm <br />0.001 m <br /> (1.7 cm17 mm <br />0.017 m <br />) +
Edinburgh New Philos. J. +
minute +
0.2 cm2 mm <br />0.002 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
black +, gray +, brown +  and tan +
angled +  and slightly rounded +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
Illustrated +  and Endemic +
0.02 cm0.2 mm <br />2.0e-4 m <br /> (0.03 cm0.3 mm <br />3.0e-4 m <br />) +
0.4 cm4 mm <br />0.004 m <br /> (?) +  and 0.6 cm6 mm <br />0.006 m <br /> (?) +
subalternate +, alternate +, whorled +  and opposite +
55 cm550 mm <br />0.55 m <br /> (70 cm700 mm <br />0.7 m <br />) +
glutinous +
climbing +  and scrambling +
sprawling +, creeping +  and prostrate +
ascending;erect +
not glaucous +, hairy +  and glabrous +
3 cm30 mm <br />0.03 m <br /> (55 cm550 mm <br />0.55 m <br />) +
0.4 cm4 mm <br />0.004 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
papillate +  and smooth +
denticulate +
Penstemon confertus var. procerus +
Penstemon procerus +
Penstemon sect. Penstemon +
species +
0.2 cm2 mm <br />0.002 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
2-ridged +  and inflated +
not constricted +
cylindric +
0.5 cm5 mm <br />0.005 m <br /> (23 cm230 mm <br />0.23 m <br />) +
straight +
gibbous +  and not spurred +
0.2 cm2 mm <br />0.002 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
cauline +  and proximal +