Potentilla arenosa

(Turczaninow) Juzepczuk in V. L. Komarov et al.

in V. L. Komarov et al., Fl. URSS 10: 137. 1941.

Basionyms: Potentilla nivea var. arenosa Turczaninow Bull. Soc. Imp. Naturalistes Moscou 16: 607. 1843
Treatment appears in FNA Volume 9. Treatment on page 200. Mentioned on page 196, 197, 199, 201, 205, 206, 208.

Plants scarcely to ± tufted. Caudex branches thick, not columnar, not sheathed with marcescent whole leaves. Stems ascending to erect, (0.3–) 0.8–2.5 (–4.5) dm, lengths (2–) 3–5 times basal leaves. Basal leaves 1.5–12 (–20) cm; petiole 1–7 (–15) cm, long hairs sparse to abundant, spreading to ± ascending, rarely loosely appressed, 1–2 (–2.5) mm, usually stiff, sometimes weak (subsp. chamissonis), verrucose, short and/or crisped hairs absent or sparse to abundant, cottony hairs absent, glands absent or sparse; leaflets separate to ± overlapping, central obovate, 1–3.5 (–4.5) × 0.5–2 (–3) cm, usually petiolulate, petiolule to 5 mm, base cuneate, margins slightly revolute, distal ± 3/4 incised ± 1/2 to midvein, teeth (2–) 3–4 (–6) per side, ± approximate to distant, surfaces dissimilar, often strongly so, abaxial white to gray, long hairs 0.5–1.8 mm, cottony-crisped hairs ± dense, adaxial green, sometimes grayish green, long hairs sparse to abundant, short-crisped hairs sparse to abundant. Cauline leaves (0–) 1–2. Inflorescence 1–7 (–15) -flowered. Pedicels 1.5–5 cm in flower, to 6 (–10) cm in fruit. Flowers: epicalyx bractlets linear-lanceolate to lanceolate-elliptic, 2–5 (–7) × 0.4–1.2 (–1.5) mm, 1/4–1/2 as wide as sepals, margins usually flat, red-glands absent or sparse and inconspicuous; hypanthium 3–5 mm diam.; sepals 3–6 (–8) mm, apex acute; petals 4–7 (–10) × 4–7 (–9) mm, ± longer than sepals; filaments 0.8–1 mm, anthers 0.4 mm; carpels 28–40, apical hairs absent, styles conic-columnar, strongly papillate-swollen in proximal 1/5–1/3, 1–1.5 mm. Achenes 1.1 mm.

Distribution

V9 301-distribution-map.jpg

Alta., B.C., Man., N.W.T., Nunavut, Que., Sask., Yukon, Alaska, Eurasia

Discussion

Subspecies 2 (2 in the flora).

The name Potentilla arenosa is now used for most arctic and subarctic plants previously treated as P. hookeriana or P. nivea subsp. hookeriana (Lehmann) Hiitonen. As noted by J. Soják (1986), the type of P. hookeriana has quinate leaves; that name is now restricted to a Rocky Mountain species in sect. Rubricaules. The arctic and subarctic material was briefly (1989–1999) called P. nivea, as discussed under that species. Because the type of P. nivea var. arenosa and other northern Asian specimens correspond closely to the North American plants, the name P. arenosa is assigned here.

The two subspecies differ only in one character, the petiole hairs, but are largely allopatric. Subspecies arenosa occurs in western and northern Greenland, northern North America (very northern in the east), and northern Asia (and perhaps northeasternmost European Russia); subsp. chamissonis occurs in southern Greenland, northeastern North America (more southern than subsp. arenosa), and northern Europe at least east to the Urals.

Selected References

None.

Key

1 Petioles with common to abundant short and/or stiff crisped hairs in addition to long verrucose hairs. Potentilla arenosa subsp. arenosa
1 Petioles with sparse or no short and/or soft crisped hairs in addition to long verrucose hairs. Potentilla arenosa subsp. chamissonis

"dm" is not declared as a valid unit of measurement for this property.

... more about "Potentilla arenosa"
reddish +, yellowish white +, gray +  and white +
cottony-crisped +
reddish +, yellowish white +, gray +  and white +
cottony-crisped +
white +  and gray +
cottony-crisped +
rugose +  and smooth +
grayish green +  and green +
0.4 cm4 mm <br />0.004 m <br /> (?) +
Reidar Elven +, David F. Murray +  and Barbara Ertter +
(Turczaninow) Juzepczuk in V. L. Komarov et al. +
compound +  and simple +
12 cm120 mm <br />0.12 m <br /> (20 cm200 mm <br />0.2 m <br />) +
1.5 cm15 mm <br />0.015 m <br /> (12 cm120 mm <br />0.12 m <br />) +
cuneate +
Potentilla nivea var. arenosa +
25 cm250 mm <br />0.25 m <br /> (30 cm300 mm <br />0.3 m <br />) +
less cordate or reniform +  and narrowly elliptic +
0.5 cm5 mm <br />0.005 m <br /> (25 cm250 mm <br />0.25 m <br />) +
coriaceous +
reduced +
free +  and distinct +
hairy +  and glabrous +
3.5 cm35 mm <br />0.035 m <br /> (4.5 cm45 mm <br />0.045 m <br />) +
2 cm20 mm <br />0.02 m <br /> (3 cm30 mm <br />0.03 m <br />) +
1 cm10 mm <br />0.01 m <br /> (3.5 cm35 mm <br />0.035 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (2 cm20 mm <br />0.02 m <br />) +
Alta. +, B.C. +, Man. +, N.W.T. +, Nunavut +, Que. +, Sask. +, Yukon +, Alaska +  and Eurasia +
not +  and aggregated +
0.5 cm5 mm <br />0.005 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
0.12 cm1.2 mm <br />0.0012 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
linear-lanceolate +  and lanceolate-elliptic +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.12 cm1.2 mm <br />0.0012 m <br />) +
0.08 cm0.8 mm <br />8.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
0.8 cm8 mm <br />0.008 m <br /> (2 cm20 mm <br />0.02 m <br />) +
unisexual +  and bisexual +
not +  and aggregated +
deciduous +
on style scar +, hairy +  and glabrous +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.26 cm2.6 mm <br />0.0026 m <br />) +
crisped +
stoloniferous +  and short-rhizomatous +
0.2 cm2 mm <br />0.002 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
straight +
weak +  and stiff +
spreading +  and more or less ascending +
sparse +  and abundant +
verrucose +
short-crisped +, cottony-crisped +  and crisped +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.18 cm1.8 mm <br />0.0018 m <br />) +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
0.7 cm7 mm <br />0.007 m <br /> (1 cm10 mm <br />0.01 m <br />) +
0.3 cm3 mm <br />0.003 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
1-7(-15)-flowered +
crassinucellate +
bipinnate +, odd-pinnate +, subpalmate +, palmate +  and ternate +
opposite +  and alternate +
marcescent +
well-developed +
persistent +
cauline +  and basal +
reduced +
separate +  and more or less overlapping +
obtriangular +, suborbiculate +, elliptic +  and obovate +
linear +  and oblanceolate +
palmate +  and pinnate +
1/2 +  and 3/4 +
inferior +  and superior +
clustered +, biseriate +  and superposed +
6 cm60 mm <br />0.06 m <br /> (10 cm100 mm <br />0.1 m <br />) +
0 cm0 mm <br />0 m <br /> (6 cm60 mm <br />0.06 m <br />) +
0.7 cm7 mm <br />0.007 m <br /> (1 cm10 mm <br />0.01 m <br />) +
1 cm10 mm <br />0.01 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
0.7 cm7 mm <br />0.007 m <br /> (0.9 cm9 mm <br />0.009 m <br />) +
free +  and distinct +
0.4 cm4 mm <br />0.004 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
more or less longer +
0.4 cm4 mm <br />0.004 m <br /> (1 cm10 mm <br />0.01 m <br />) +
0.4 cm4 mm <br />0.004 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
7 cm70 mm <br />0.07 m <br /> (15 cm150 mm <br />0.15 m <br />) +
1 cm10 mm <br />0.01 m <br /> (7 cm70 mm <br />0.07 m <br />) +
0 cm0 mm <br />0 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
adnate +, free +, connate +  and distinct +
palmate +  and ternate +
12 cm120 mm <br />0.12 m <br /> (20 cm200 mm <br />0.2 m <br />) +
1 cm10 mm <br />0.01 m <br /> (12 cm120 mm <br />0.12 m <br />) +
in V. L. Komarov et al., Fl. URSS +
inconspicuous +
not arillate +
0.6 cm6 mm <br />0.006 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
persistent +
free +  and distinct +
erect +  and spreading +
lanceolate +  and broadly ovate or deltate +
0.3 cm3 mm <br />0.003 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
free +  and distinct +
not rooting +
reddish +  and green +
(2-)3-5 times basal leaves +
ascending +  and erect +
1 +  and many +
not flagelliform +
palmate +  and pinnate +
linear;ovate +
0.12 cm1.2 mm <br />0.0012 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
deciduous +
distinct +
proximal +  and subapical +
papillate-swollen +  and conic-columnar +
rough-thickened +
0.07 cm0.7 mm <br />7.0e-4 m <br /> (0.12 cm1.2 mm <br />0.0012 m <br />) +
not fleshy-thickened +
Potentilla arenosa +
Potentilla sect. Niveae +
species +
3 +  and 4 +
inconspicuous +
enlarged +
biennial +, annual +  and perennial +