Potentilla sect. Pentaphylloides

Tausch

Hort. Canal., sub P. ornithopoda. 1823.

Synonyms: Undefined sect. Argentina Hill Potentilla sect. Anserina (Gaudin) Pfeiffer Potentilla ser. Anserina (Gaudin) T. T. Yu & C. L. Li Potentilla subg. Argentina (Hill) Jepson
Treatment appears in FNA Volume 9. Treatment on page 126. Mentioned on page 121, 124, 127.

Perennials, openly matted, stoloniferous [to ± tufted, not stoloniferous]; taproots not fleshy-thickened, often absent and replaced by ± thickened lateral roots; vestiture primarily of long and crisped-cottony hairs, glands absent or sparse, not red. Stems prostrate [to erect], flagelliform [or not], rooting at nodes [or not], lateral to ± persistent basal rosettes, 1–10+ dm, lengths (0.5–) 1–5+ times basal leaves. Leaves: basal not or ± 2-ranked; cauline (between flowering and/or rosette-forming nodes) 0–1; primary leaves pinnate (distal leaflets sometimes confluent), (1–) 3–50 (–75) cm; petiole: long hairs absent or ascending to weakly appressed, sometimes spreading, soft to weak, glands absent or very sparse; primary leaflets (5–) 7–21 (–31 [–51]) plus additional small leaflets interspersed, on distal (1/2–) 2/3–4/5+ of leaf axis, overlapping or not, elliptic to obovate, margins revolute or nearly flat, distal (1/2–) 2/3–3/4+ evenly incised 1/4–1/2 to midvein, teeth (2–) 5–12 (–16) per side, surfaces similar to strongly dissimilar, abaxial usually white, sometimes green, cottony (/crisped) hairs usually dense, sometimes sparse or absent, adaxial green to white, not glaucous, long hairs absent or weak to soft. Inflorescences solitary flowers at stolon nodes [± cymose]. Pedicels straight in fruit, (1–) 2–45 (–55) cm. Flowers 5 (–7) -merous; hypanthium 3–7 mm diam.; petals bright-yellow, elliptic to broadly elliptic, (4–) 5–15 (–20) mm, longer than sepals, apex rounded or slightly retuse; stamens [5–] 20 (–30); styles ± lateral [subapical or subbasal], narrowly columnar-filiform, not papillate-swollen proximally, 1.5–2.5 mm. Achenes slightly rugose and papillate.

Distribution

North America, Mexico, South America (Argentina), South America (Chile), Eurasia, Pacific Islands (New Guinea), Pacific Islands (New Zealand), s Australia

Discussion

Species ca. 50 (1 in the flora).

The use here of sect. Pentaphylloides is contrary to the argument by J. Soják (2007) that the use of Pentaphylloides Tournefort by G. R. Boehmer (in C. G. Ludwig 1760) qualified as an unranked infrageneric taxon, making sect. Pentaphylloides Tausch a later homonym. In that Boehmer was citing synonyms, not infrataxa, sect. Pentaphylloides Tausch is the correct name for this section. The names sect. Chenopotentilla Focke and sect. Leptostylae (Th. Wolf) Guşuleac are superfluous and illegitimate.

Potentilla anserina is the atypical and only representative in North America of a primarily southeastern Asian section. Although provisionally retained within Potentilla, the morphological distinctiveness of the section has led to the recognition of Argentina as a segregate genus (for example, P. A. Rydberg 1908d; Á Löve and D. Löve 1975b; B. C. Johnston 1985; J. Soják 2010). Molecular analyses (C. Dobeš and J. Paule 2010; M. H. Töpel et al. 2011) have confirmed the section is a strongly supported monophyletic clade, albeit including some other Asian species treated by J. Soják (1994) as the segregate genera Piletophyllum (Soják) Soják and Tylosperma Botschantzev. Chloroplast markers (Dobeš and Paule; Töpel et al.) place this clade sister to all other Potentilleae; nuclear markers (Töpel et al.) indicate instead a sister relation to Fragariinae.

Selected References

None.

"thin" is not a number."(" is not declared as a valid unit of measurement for this property."dm" is not declared as a valid unit of measurement for this property."dm" is not declared as a valid unit of measurement for this property.

papillate +  and rugose +
green +  and white +
retuse;rounded +
Reidar Elven +  and David F. Murray +
Tausch +
compound +  and simple +
25 cm250 mm <br />0.25 m <br /> (30 cm300 mm <br />0.3 m <br />) +
less cordate or reniform +  and narrowly elliptic +
0.5 cm5 mm <br />0.005 m <br /> (25 cm250 mm <br />0.25 m <br />) +
coriaceous +
reduced +
free +  and distinct +
hairy +  and glabrous +
North America +, Mexico +, South America (Argentina) +, South America (Chile) +, Eurasia +, Pacific Islands (New Guinea) +, Pacific Islands (New Zealand) +  and s Australia +
not +  and aggregated +
0.8 cm8 mm <br />0.008 m <br /> (2 cm20 mm <br />0.02 m <br />) +
unisexual +  and bisexual +
not +  and aggregated +
deciduous +
on style scar +, hairy +  and glabrous +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.26 cm2.6 mm <br />0.0026 m <br />) +
stoloniferous +  and short-rhizomatous +
straight +
cottony +  and crisped-cottony +
sparse +  and ascending to weakly appressed +
crisped +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
0.7 cm7 mm <br />0.007 m <br /> (1 cm10 mm <br />0.01 m <br />) +
0.3 cm3 mm <br />0.003 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
0.5mm;2.5mm +
turbinate +, patelliform +  and cupulate +
racemiform open +  and more or less congested +
crassinucellate +
bipinnate +, odd-pinnate +, subpalmate +, palmate +  and ternate +
opposite +  and alternate +
marcescent +
well-developed +
persistent +
cauline +  and basal +
reduced +
not +  and overlapping +
elliptic +  and obovate +
linear +  and oblanceolate +
palmate +  and pinnate +
1/4 (?) +  and 1/2 (?) +
incised +, flat +  and revolute +
inferior +  and superior +
clustered +, biseriate +  and superposed +
45 cm450 mm <br />0.45 m <br /> (55 cm550 mm <br />0.55 m <br />) +
2 cm20 mm <br />0.02 m <br /> (45 cm450 mm <br />0.45 m <br />) +
elliptic +  and broadly elliptic +
1.5 cm15 mm <br />0.015 m <br /> (2 cm20 mm <br />0.02 m <br />) +
bright-yellow +
free +  and distinct +
commonly obcordate;nearly round +
0.5 cm5 mm <br />0.005 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
adnate +, free +, connate +  and distinct +
50 cm500 mm <br />0.5 m <br /> (75 cm750 mm <br />0.75 m <br />) +
3 cm30 mm <br />0.03 m <br /> (50 cm500 mm <br />0.5 m <br />) +
Hort. Canal., sub P. ornithopoda. +
not arillate +
persistent +
free +  and distinct +
erect +  and spreading +
lanceolate +  and broadly ovate or deltate +
free +  and distinct +
reddish +  and green +
prostrate +  and erect +
lateral;more or less persistent basal +
1 +  and many +
flagelliform +
palmate +  and pinnate +
linear;ovate +
deciduous +
distinct +
papillate-swollen +  and columnar-filiform +
rough-thickened +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
Undefined sect. Argentina +, Potentilla sect. Anserina +, Potentilla ser. +  and Potentilla subg. Argentina +
not fleshy-thickened +
Potentilla sect. Pentaphylloides +
Potentilla +
section +
5 (?) +  and 12 (?) +
inconspicuous +
enlarged +
biennial +, annual +  and perennial +
herb +  and matted +