Potentilla subvahliana

Jurtzev in A. I. Tolmatchew

in A. I. Tolmatchew, Fl. Arct. URSS 9(1): 319. 1984.

Illustrated
Treatment appears in FNA Volume 9. Treatment on page 203. Mentioned on page 196, 198, 202, 205, 210.

Plants usually cushion-forming. Caudex branches stout, columnar, sheathed with marcescent whole leaves. Stems erect, 0.2–0.6 (–1.1) dm, lengths 2–4 times basal leaves. Basal leaves 0.5–2.5 (–3) cm; petiole 0.3–1.5 (–2) cm, long hairs common to abundant, spreading to ascending, 1 (–2) mm, ± weak, rarely stiff, smooth, crisped/short-cottony hairs usually absent, sometimes sparse, glands sparse to common; leaflets separate to slightly overlapping, central obovate, 0.5–1.2 (–1.5) × 0.4–0.9 (–1) cm, subsessile to short-petiolulate, base cuneate, margins revolute, distal 1/2 incised 1/2–3/4 to midvein, teeth (1–) 2–3 (–4) per side, ± distant, surfaces ± dissimilar, abaxial grayish or yellowish white, long hairs 0.5–1 mm, cottony-crisped hairs ± dense (often obscured by long hairs), adaxial dark green to greenish gray, long hairs ± abundant, other hairs absent. Cauline leaves (0–) 1 (–2). Inflorescences 1 (–2) -flowered. Pedicels 1–2 (–3) cm in flower, to 5.5 cm in fruit. Flowers: epicalyx bractlets lanceolate to elliptic-ovate, rarely linear, 3–6 (–7) × 0.8–2 (–2.3) mm, (1/2–) 2/3 to nearly as wide as sepals, margins flat or ± revolute, red-glands absent; hypanthium 2.5–4 mm diam.; sepals 3–5 (–6) mm, apex obtuse to subacute; petals 4–8 (–9) × 4–9 mm, longer than sepals; filaments 1–1.3 mm, anthers 0.3–0.5 mm; carpels 25–35, apical hairs absent, styles conic-columnar, ± papillate-swollen on less than proximal 1/5, 0.9–1.1 mm. Achenes 1.2–1.6 mm. 2n = 28 (Russian Far East).


Phenology: Flowering late spring to summer.
Habitat: Rocky alpine heaths, rock outcrops and crevices, scree and talus, dry tundra, coastal bluffs, stabilized sand dunes, usually on calcareous bedrock
Elevation: 0–1700 m

Distribution

V9 308-distribution-map.jpg

Greenland, Alta., B.C., N.W.T., Nunavut, Que., Yukon, Alaska, e Asia (Russian Far East)

Discussion

The major part of the range assigned to Potentilla vahliana by E. Hultén (1968) belongs to P. subvahliana.

Two morphologic types are present within what is accepted here as Potentilla subvahliana. Plants corresponding to the type of P. subvahliana (Wrangel Island) are widespread throughout northeastern Asia (Chukotka) and arctic North America to northwestern Greenland. Some plants in alpine Alaska, Yukon, and east to Amundsen Gulf, Nunavut, are redder, more densely columnar, and less hairy, with smaller leaves having fewer and narrower lobes, more slender one-flowered stems, narrow and entire bracts, narrower sepals, and much narrower epicalyx bractlets. Potentilla subvahliana is tetraploid, perhaps an allopolyploid, and could consist of lineages from different parental combinations.

Selected References

None.

Lower Taxa

None.

"dm" is not declared as a valid unit of measurement for this property.

... more about "Potentilla subvahliana"
reddish +, yellowish white +, gray +  and white +
cottony-crisped +
reddish +, yellowish white +, gray +  and white +
cottony-crisped +
yellowish white +  and grayish +
cottony-crisped +
rugose +  and smooth +
1.2mm +  and 1.6mm +
dark green +  and greenish gray +
0.03 cm0.3 mm <br />3.0e-4 m <br /> (0.05 cm0.5 mm <br />5.0e-4 m <br />) +
Reidar Elven +, David F. Murray +  and Barbara Ertter +
Jurtzev in A. I. Tolmatchew +
compound +  and simple +
2.5 cm25 mm <br />0.025 m <br /> (3 cm30 mm <br />0.03 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
cuneate +
Niveae +
25 cm250 mm <br />0.25 m <br /> (30 cm300 mm <br />0.3 m <br />) +
less cordate or reniform +  and narrowly elliptic +
0.5 cm5 mm <br />0.005 m <br /> (25 cm250 mm <br />0.25 m <br />) +
coriaceous +
reduced +
free +  and distinct +
hairy +  and glabrous +
subsessile +  and short-petiolulate +
1.2 cm12 mm <br />0.012 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
0.9 cm9 mm <br />0.009 m <br /> (1 cm10 mm <br />0.01 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
0.4 cm4 mm <br />0.004 m <br /> (0.9 cm9 mm <br />0.009 m <br />) +
Greenland +, Alta. +, B.C. +, N.W.T. +, Nunavut +, Que. +, Yukon +, Alaska +  and e Asia (Russian Far East) +
not +  and aggregated +
0–1700 m +
0.6 cm6 mm <br />0.006 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (0.23 cm2.3 mm <br />0.0023 m <br />) +
0.3 cm3 mm <br />0.003 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
lanceolate +  and elliptic-ovate +
0.08 cm0.8 mm <br />8.0e-4 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
0.1 cm1 mm <br />0.001 m <br /> (0.13 cm1.3 mm <br />0.0013 m <br />) +
0.8 cm8 mm <br />0.008 m <br /> (2 cm20 mm <br />0.02 m <br />) +
unisexual +  and bisexual +
not +  and aggregated +
deciduous +
on style scar +, hairy +  and glabrous +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.26 cm2.6 mm <br />0.0026 m <br />) +
crisped +
Rocky alpine heaths, rock outcrops and crevices, scree and talus, dry tundra, coastal bluffs, stabilized sand dunes, usually on calcareous bedrock +
stoloniferous +  and short-rhizomatous +
0.1 cm1 mm <br />0.001 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
straight +
stiff +  and weak +
spreading +  and ascending +
abundant +  and common +
cottony-crisped +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
0.7 cm7 mm <br />0.007 m <br /> (1 cm10 mm <br />0.01 m <br />) +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
crassinucellate +
bipinnate +, odd-pinnate +, subpalmate +, palmate +  and ternate +
opposite +  and alternate +
marcescent +
well-developed +
persistent +
cauline +  and basal +
reduced +
separate +  and slightly overlapping +
obtriangular +, suborbiculate +, elliptic +  and obovate +
linear +  and oblanceolate +
palmate +  and pinnate +
1/2 +  and 3/4 +
revolute +, flat +  and incised +
inferior +  and superior +
clustered +, biseriate +  and superposed +
2 cm20 mm <br />0.02 m <br /> (3 cm30 mm <br />0.03 m <br />) +
0 cm0 mm <br />0 m <br /> (5.5 cm55 mm <br />0.055 m <br />) +
0.8 cm8 mm <br />0.008 m <br /> (0.9 cm9 mm <br />0.009 m <br />) +
1 cm10 mm <br />0.01 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
free +  and distinct +
0.4 cm4 mm <br />0.004 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
0.4 cm4 mm <br />0.004 m <br /> (1 cm10 mm <br />0.01 m <br />) +
0.4 cm4 mm <br />0.004 m <br /> (0.9 cm9 mm <br />0.009 m <br />) +
1.5 cm15 mm <br />0.015 m <br /> (2 cm20 mm <br />0.02 m <br />) +
0.3 cm3 mm <br />0.003 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
Flowering late spring to summer. +
adnate +, free +, connate +  and distinct +
palmate +  and ternate +
12 cm120 mm <br />0.12 m <br /> (20 cm200 mm <br />0.2 m <br />) +
1 cm10 mm <br />0.01 m <br /> (12 cm120 mm <br />0.12 m <br />) +
in A. I. Tolmatchew, Fl. Arct. URSS +
not arillate +
0.5 cm5 mm <br />0.005 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
persistent +
free +  and distinct +
erect +  and spreading +
lanceolate +  and broadly ovate or deltate +
0.3 cm3 mm <br />0.003 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
Illustrated +
free +  and distinct +
not rooting +
reddish +  and green +
2-4 times basal leaves +
1 +  and many +
not flagelliform +
palmate +  and pinnate +
linear;ovate +
0.12 cm1.2 mm <br />0.0012 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
deciduous +
distinct +
proximal +  and subapical +
papillate-swollen +  and conic-columnar +
rough-thickened +
0.09 cm0.9 mm <br />9.0e-4 m <br /> (0.11 cm1.1 mm <br />0.0011 m <br />) +
not fleshy-thickened +
Potentilla subvahliana +
Potentilla sect. Niveae +
species +
2 +  and 3 +
inconspicuous +
enlarged +
biennial +, annual +  and perennial +
plant +  and cushion-forming +