Poterium sanguisorba var. polygamum

(Waldstein & Kitaibel) Visiani

Fl. Dalmat. 3: 255. 1851.

Common names: Salad or fodder burnet burnet bloodwort sanguisorbe mineure
IllustratedIntroduced
Basionym: Poterium polygamum Waldstein & Kitaibel Descr. Icon. Pl. Hung. 2: 217, plate 198. 1805
Synonyms: P. balearicum (Nyman) Bourgeau ex Porta P. sanguisorba subsp. muricatum Bonnier & Layens Sanguisorba minor subsp. balearica (Nyman) Muñoz Garmendia & C. Navarro S. minor subsp. muricata (Bonnier & Layens) Briquet
Treatment appears in FNA Volume 9. Treatment on page 318. Mentioned on page 319.

Leaves: stipules of basal leaves simple, lanceolate, margins entire, those of cauline lanceolate to ovate, coarsely serrate; petiole longer than blade in basal and proximal cauline leaves, to shorter than proximal rachis segment in distal cauline ones; leaflets 2–12 pairs, proximally alternate, subopposite, or opposite, distally subopposite or opposite, margins coarsely serrate, surfaces glabrous or sparsely pilose. Spikes subglobose to ovoid, 0.3–2.5 × 0.3–2 cm; peduncles glabrous or sparsely loosely villous; bracteoles ovate, equal to hypanthia, ciliate. Flowers generally gynomonoecious, bisexual flowers proximal and pistillate distal (rarely some spikes completely pistillate or completely bisexual); hypanthium broadly ellipsoid; sepals green to purple, margins scarious, apex acute to apiculate with tufts of trichomes; filaments filiform, equal to sepals. Achenes enclosed within broadly ellipsoid hypanthia 4 mm, 4-sided, angles winged, surfaces irregularly reticulately ridged, ridges sometimes also muricate. 2n = 28, 56.


Phenology: Flowering May–Aug; fruiting Jul–Oct.
Habitat: Open disturbed areas, pastures, roadsides, woodland edges, along railroad rights-of-way, some preference for calcareous soil
Elevation: 0–2700 m

Distribution

V9 519-distribution-map.jpg

Introduced; B.C., N.S., Ont., Que., Ala., Ariz., Calif., Colo., Conn., Del., Idaho, Ill., Ind., Md., Mass., Mich., Mont., Nev., N.J., N.Mex., N.Y., N.C., Ohio, Oreg., Pa., R.I., Tenn., Tex., Utah, Vt., Va., Wash., W.Va., Wyo., Europe, w Asia, n Africa, also in Australia

Discussion

All North American plants of Poterium belong to var. polygamum. This taxon has also been reported for New Brunswick; this is unconfirmed, but certainly plausible. Reports for North America of Sanguisorba minor or P. sanguisorba refer to that in a broad sense, not distinguishing the varieties or subspecies variously recognized, and do not imply the occurrence of the typical infrataxon in North America. It is possible that the typical variety (or other infrataxa, as defined in the European literature) is present in North America. The other infrataxa differ primarily in details of the ornamentation of the hypanthium, with either more or less pronounced wings, differences in the roundedness or acuteness of the wings, and variations in the reticulate patterns between the wings; refer to the European literature to identify material suspected of belonging to other infrataxa.

Selected References

None.

Lower Taxa

None.
Alan S. Weakley +
(Waldstein & Kitaibel) Visiani +
compound +  and simple +
opposite +  and alternate +
Poterium polygamum +
15 cm150 mm <br />0.15 m <br /> (35 cm350 mm <br />0.35 m <br />) +
oblanceolate +  and obovate +
1 cm10 mm <br />0.01 m <br /> (15 cm150 mm <br />0.15 m <br />) +
free +  and distinct +
Salad or fodder burnet +, burnet bloodwort +  and sanguisorbe mineure +
B.C. +, N.S. +, Ont. +, Que. +, Ala. +, Ariz. +, Calif. +, Colo. +, Conn. +, Del. +, Idaho +, Ill. +, Ind. +, Md. +, Mass. +, Mich. +, Mont. +, Nev. +, N.J. +, N.Mex. +, N.Y. +, N.C. +, Ohio +, Oreg. +, Pa. +, R.I. +, Tenn. +, Tex. +, Utah +, Vt. +, Va. +, Wash. +, W.Va. +, Wyo. +, Europe +, w Asia +, n Africa +  and also in Australia +
not +  and aggregated +
0–2700 m +
0.2 cm2 mm <br />0.002 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
bisexual +  and gynomonoecious +
not +  and aggregated +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
Open disturbed areas, pastures, roadsides, woodland edges, along railroad rights-of-way, some preference for calcareous soil +
gland-tipped +, unicellular +  and multicellular +
acicular +
tan +  and brown +
glabrous +  and hairy +
0.2 cm2 mm <br />0.002 m <br /> (0.9 cm9 mm <br />0.009 m <br />) +
[15-]20-100+-flowered +
crassinucellate +
deciduous +
opposite +, subopposite +  and alternate +
obovate +, orbiculate +  and elliptic +
serrate +  and entire +
lanceolate +  and ovate +
inferior +  and superior +
clustered +, biseriate +  and superposed +
villous +  and glabrous +
0 (?) +  and 4 (?) +
free +  and distinct +
5 +  and 4 +
Flowering May–Aug +  and fruiting Jul–Oct. +
adnate +, free +, connate +  and distinct +
Fl. Dalmat. +
muricate +
not arillate +
green +  and purple +
persistent +
distinct +
spreading +  and slightly ascending +
ovate +  and elliptic +
Illustrated +  and Introduced +
0.3 cm3 mm <br />0.003 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
pubescent +  and glabrous +
subglobose +  and ovoid +
0.3 cm3 mm <br />0.003 m <br /> (2 cm20 mm <br />0.02 m <br />) +
free +  and distinct +
shorter or longer +
branched +  and simple +
ascending +  and erect +
1 +  and 10 +
deciduous +
distinct +
basal +, lateral +, subterminal +  and terminal +
not elongate +
pilose +  and glabrous +
P. balearicum +, P. sanguisorba subsp. muricatum +, Sanguisorba minor subsp. balearica +  and S. minor subsp. muricata +
Poterium sanguisorba var. polygamum +
Poterium sanguisorba +
variety +
inconspicuous +
enlarged +  and small +