Purshia subintegra

(Kearney) Henrickson

Phytologia 60: 468. 1986.

Common names: Arizona cliffrose
Conservation concernEndemic
Basionym: Cowania subintegra Kearney Madroño 7: 15. 1943
Synonyms: Purshia pinkavae Schaack
Treatment appears in FNA Volume 9. Treatment on page 342. Mentioned on page 339, 341.

Shrubs, evergreen or tardily drought-deciduous, 2–8 (–12) dm. Stems: young-stem internodes 2.5–9 (–13) mm, closely arachnoid-villous, sometimes pubescent-hirtellous, rarely stipitate-glandular; short-shoot spurs simple, 1–10 × 1–1.3 mm. Leaves: blade green adaxially, linear to oblanceolate, (3–) 4–8 (–10) [–17] × 1–1.3 (–2) [–4.2] mm, margins revolute (not obscuring abaxial surfaces), entire or with 1–2 teeth or lobes, lobes ascending, oblong, 1 (–2) mm, margins strongly revolute, apex obtuse-rounded, sometimes ± apiculate, abaxial surface densely arachnoid-villous, midvein glabrous (when exposed), adaxial hirtellous (along midvein proximally), initially closely arachnoid-villous, glabrescent, eglandular. Pedicels 1.5–6 (–12) mm. Flowers: hypanthium narrowly obconic, 3–6 × 1.5–3.2 mm, 4.5 mm diam. in fruit, arachnoid-villous, rarely stipitate-glandular; petals light yellowish cream, obovate-spatulate to spatulate, (4–) 6–8 mm; stamens (28–) 34–54; carpels (2–) 4–5 (–6). Achenes oblanceoloid, 5–6 × 1.5–2.5 mm, 13–15 ribbed, persistent style 32–45 mm, plumose hairs 1.5–3 mm. 2n = 18.


Phenology: Flowering Mar–early Jun.
Habitat: Whitish, lacustrine-derived limy soil of Miocene-Pliocene age
Elevation: 600–1100 m

Discussion

Of conservation concern.

Purshia subintegra is known from four populations in Arizona: Mojave County (north of Burro Creek); Maricopa and adjacent Yavapai counties (Tonto Basin, western side of Horseshoe Reservoir on Verde River); Yavapai County (Verde Valley, south of Cottonwood, the largest population); and Graham County (San Carlos Basin northwest of Bylas). The first three populations consist of plants with both entire or lobed leaves (on individual plants) and some stipitate glands on pedicels and hypanthia. The Bylas population consists mainly of plants with mostly entire leaves and without stipitate glands, although some collections from that population have stipitate glands and others have lobed long-shoot leaves. The lobed leaves and stipitate glands have been considered markers of introgression with P. stansburyana, which has been confirmed by morphometric studies (F. W. Reichenbacher 1994) and by molecular markers (S. E. Travis et al. 2008) that provide evidence of introgression with P. stansburyana in all but the Bylas population.

Because of this variation and introgression, C. G. Schaack (1987) lectotypified Purshia subintegra on a specimen with lobed leaves, relegating that name to hybrid status. He named a new taxon (P. pinkavae) from the population near Bylas that mostly (but not entirely) lacks stipitate glands and lobed leaves; this seems unnecessary, as edaphic and morphologic continuity exists among known populations of P. subintegra, and all four populations share at least some traits with P. stansburyana as well as with P. ericifolia.

Selected References

None.

Lower Taxa

None.

"-6×1.5-3.2" is not declared as a valid unit of measurement for this property.

... more about "Purshia subintegra"
arachnoid-villous +
0.5 cm5 mm <br />0.005 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
oblanceoloid +
1.5mm +  and 2.5mm +
glabrescent +, arachnoid-villous +  and hirtellous +
James Henrickson +
(Kearney) Henrickson +
gray +  and cinnamon brown +
weathering +
compound +  and simple +
opposite +  and alternate +
cuneate +  and clawed +
Cowania subintegra +
0.8 cm8 mm <br />0.008 m <br /> (1 cm10 mm <br />0.01 m <br />) +
1.5 cm15 mm <br />0.015 m <br /> (1.9 cm19 mm <br />0.019 m <br />) +
0.13 cm1.3 mm <br />0.0013 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
0.4 cm4 mm <br />0.004 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
linear +  and oblanceolate +
0.3 cm3 mm <br />0.003 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
1mm +  and 1.3mm +
free +  and distinct +
Arizona cliffrose +
not +  and aggregated +
600–1100 m +
nectariferous +
glabrous +, not +, stipitate-glandular +  and villous +
1 cm10 mm <br />0.01 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
unisexual +  and bisexual +
not +  and aggregated +
oblanceoloid +  and obovoid-fusiform +
0.4 cm4 mm <br />0.004 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
Whitish, lacustrine-derived limy soil of Miocene-Pliocene age +
plumose +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
stipitate-glandular +  and arachnoid-villous +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.65 cm6.5 mm <br />0.0065 m <br />) +
crassinucellate +
drought-deciduous +  and persistent +
immature +  and abscising +
0.1 cm1 mm <br />0.001 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
with 1-2 teeth or lobes +  and entire +
toothed +  and lobed +
inferior +  and superior +
clustered +, biseriate +  and superposed +
0.6 cm6 mm <br />0.006 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
0 (?) +  and 4 (?) +
0.4 cm4 mm <br />0.004 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
free +  and distinct +
0.6 cm6 mm <br />0.006 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
Flowering Mar–early Jun. +
adnate +, free +, connate +  and distinct +
not arillate +
persistent +
free +  and distinct +
spreading-reflexing +  and spreading-reflexed +
ovate +  and oblong-ovate +
0.1 cm1 mm <br />0.001 m <br /> (1 cm10 mm <br />0.01 m <br />) +
0.1 cm1 mm <br />0.001 m <br /> (0.13 cm1.3 mm <br />0.0013 m <br />) +
Conservation concern +  and Endemic +
free +  and distinct +
3 +  and 20 +
adnate +  and free +
persistent +
distinct +
basal +, lateral +, subterminal +  and terminal +
elongate +
3.2 cm32 mm <br />0.032 m <br /> (4.5 cm45 mm <br />0.045 m <br />) +
Purshia pinkavae +
Purshia subintegra +
species +
1 +  and 2 +
inconspicuous +
hemispheric +
drought-deciduous +  and evergreen +
0.9 cm9 mm <br />0.009 m <br /> (1.3 cm13 mm <br />0.013 m <br />) +
stipitate-glandular +, pubescent-hirtellous +  and arachnoid-villous +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.9 cm9 mm <br />0.009 m <br />) +