Pyrola picta

Smith in A. Rees

in A. Rees, Cycl., 29: Pyrola no. 8. 1814 ,.

Common names: White-vein wintergreen
IllustratedEndemic
Synonyms: Pyrola aphylla Smith Pyrola dentata Smith Pyrola dentata var. integra A. Gray Pyrola picta subsp. dentata (Smith) Piper Pyrola picta var. dentata (Smith) Dorn Pyrola picta subsp. integra (A. Gray) Piper
Treatment appears in FNA Volume 8. Treatment on page 383. Mentioned on page 378, 379, 382, 384.

Plants rhizomatous, (0.5–) 1.5–4.6 dm. Leaves sometimes highly reduced or absent; petiole 4–25 mm, channeled adaxially, glabrous; blade usually maculate, sometimes not maculate, dull and light green to purplish abaxially, dull or shiny and green to dark green, usually with white tissue bordering larger veins adaxially, ovate or ovate-elliptic to oblanceolate or spatulate, 8–80 × 4–49 mm, coriaceous, base obtuse to attenuate or decurrent, margins entire or denticulate to coarsely denticulate, apex obtuse to acute. Inflorescences 1–2 (–3), 4–24-flowered; peduncular bracts 1–5, subulate to narrowly lanceolate, 3–9 × 1–2 mm, membranous, margins entire or erose-denticulate; inflorescence bracts subulate to ovatelanceolate, usually shorter than or, rarely, longer than subtended pedicels, 3.5–6 × 2–3 mm, herbaceous. Pedicels 3–18 mm. Flowers: calyx lobes appressed or spreading in fruit, green with margins hyaline to white or pinkish, widely deltate to widely ovate, 1.1–3.9 × 1.1–2.4 mm, margins entire or erose-denticulate, apices acute; petals greenish white, white, pink, or reddish, obovate, 4.2–11.2 × 3.1–7.1 mm, margins entire or obscurely denticulate; stamens 5–9 mm; filament base 0.5–0.8 mm wide; anthers 1.8–5.5 mm, apiculations absent or 0.1–0.2 mm, thecae creamy white to pink, tubules orange, 0.4–0.8 mm, gradually narrowed from thecae, lateral walls usually touching for most of their lengths, pores 0.1–0.3 × 0.1–0.2 mm; ovary smooth; style exserted, 4–9 mm; stigma (0.8–) 1–1.6 mm wide, lobes erect. Capsules depressed-globose, 3–6 × 6–9 mm. 2n = 46.


Phenology: Flowering Jun–Aug.
Habitat: Dry, coniferous forests
Elevation: 400-3000 m

Distribution

V8 724-distribution-map.gif

Alta., B.C., Ariz., Calif., Colo., Idaho, Mont., Nev., N.Mex., Oreg., S.Dak., Utah, Wash., Wyo.

Discussion

E. Haber (1987) concluded that Pyrola picta, P. aphylla, and P. dentata are morphotypes of a single, highly variable species, a finding consistent with seed morphology data compiled by Hiroshi Takahashi (1993). Leafless scapes frequently are found attached to rhizomes bearing leafy shoots (W. H. Camp 1940; Haber 1987). Putative hybrids between P. picta and P. chlorantha have been reported at three locations in the western United States (Haber 1993). Cladistic analyses of molecular and morphologic data suggest that P. picta is sister to P. chlorantha (J. V. Freudenstein 1999b), which also occasionally is leafless.

Selected References

None.

Lower Taxa

None.

"broad" is not a number.

... more about "Pyrola picta"
0.18 cm1.8 mm <br />0.0018 m <br /> (0.55 cm5.5 mm <br />0.0055 m <br />) +
Acute (?) +  and Obtuse (?) +
Craig C. Freeman +
Smith in A. Rees +
obtuse;attenuate or decurrent +
green +  and dark green +
0.8 cm8 mm <br />0.008 m <br /> (8 cm80 mm <br />0.08 m <br />) +
shiny +  and dull +
ovate-elliptic;oblanceolate or spatulate +
coriaceous +
4mm;49mm +
0.11 cm1.1 mm <br />0.0011 m <br /> (0.39 cm3.9 mm <br />0.0039 m <br />) +
spreading +  and appressed +
0.11 cm1.1 mm <br />0.0011 m <br /> (0.24 cm2.4 mm <br />0.0024 m <br />) +
0.3 cm3 mm <br />0.003 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
depressed-globose +
0.6 cm6 mm <br />0.006 m <br /> (0.9 cm9 mm <br />0.009 m <br />) +
pseudoverticillate +, in basal rosettes +  and alternate +
White-vein wintergreen +
persistent +  and deciduous +
rotate to crateriform campanulate cylindric globose or urceolate +
Alta. +, B.C. +, Ariz. +, Calif. +, Colo. +, Idaho +, Mont. +, Nev. +, N.Mex. +, Oreg. +, S.Dak. +, Utah +, Wash. +  and Wyo. +
400-3000 m +
undifferentiated +
fusiform +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.08 cm0.8 mm <br />8.0e-4 m <br />) +
nodding +  and spreading +
unisexual +  and bisexual +
loculicidal +
Dry, coniferous forests +
multicellular +
0.35 cm3.5 mm <br />0.0035 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
subulate +  and ovatelanceolate +
0.2 cm2 mm <br />0.002 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
persistent +
scale-like +
reduced +
hyaline +  and white or pinkish +
denticulate;entire;erose-denticulate;entire;erose-denticulate;entire;denticulate;coarsely denticulate +
parietal +, axile +  and placentation +
intruded-parietal +
tenuinucellate +  and unitegmic +
0.3 cm3 mm <br />0.003 m <br /> (1.8 cm18 mm <br />0.018 m <br />) +
0.3 cm3 mm <br />0.003 m <br /> (0.9 cm9 mm <br />0.009 m <br />) +
subulate +  and narrowly lanceolate +
0.1 cm1 mm <br />0.001 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
distinct +
0.42 cm4.2 mm <br />0.0042 m <br /> (1.12 cm11.2 mm <br />0.0112 m <br />) +
reduced +
not sticky +
0.31 cm3.1 mm <br />0.0031 m <br /> (0.71 cm7.1 mm <br />0.0071 m <br />) +
channeled +
0.4 cm4 mm <br />0.004 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
Flowering Jun–Aug. +
0.01 cm0.1 mm <br />1.0e-4 m <br /> (0.03 cm0.3 mm <br />3.0e-4 m <br />) +
0.01 cm0.1 mm <br />1.0e-4 m <br /> (0.02 cm0.2 mm <br />2.0e-4 m <br />) +
in A. Rees, Cycl., +
distinct +
fusiform +
Illustrated +  and Endemic +
peltate +  and capitate +
1mm;1.6mm +
straight +
expanded +
0.4 cm4 mm <br />0.004 m <br /> (0.9 cm9 mm <br />0.009 m <br />) +
Pyrola aphylla +, Pyrola dentata +, Pyrola dentata var. integra +, Pyrola picta subsp. dentata +, Pyrola picta var. dentata +  and Pyrola picta subsp. integra +
Pyrola picta +
species +
creamy white;pink +
narrowed +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.08 cm0.8 mm <br />8.0e-4 m <br />) +
evergreen +, deciduous +  and perennial +