Quercus cornelius-mulleri

Nixon & K. P. Steele

Madroño 28: 210. 1981.

Common names: Muller oak
Illustrated
Treatment appears in FNA Volume 3.

Shrubs, evergreen or subevergreen, densely branched, 1-2 (-3) m. Bark gray, scaly. Twigs gray, yellowish, or brownish, 1-1.5 mm diam., densely tomentulose, rarely glabrate. Buds dull brown, ovoid, 2.5-3 mm, glabrous except for margins of scales. Leaves: petiole 2-5 mm. Leaf-blade strongly bicolored, ovate to oblong or narrowly obovate, 15-35 ×10-20 mm, rather thick and leathery, base cuneate or attenuate-rounded, margins entire or irregularly, shallowly toothed, teeth mucronate, rarely spinose, secondary-veins 6-7 on each side, apex rounded or acute; surfaces abaxially whitish, densely covered with minute, compact, appressed, (8-) 10-14 (-16) -rayed stellate hairs less than 0.2 mm diam. (lateral fusion of rays visible under high magnification), without glandular seriate hairs, adaxially dull, light green, grayish, or yellowish green, with scattered, appressed-stellate hairs to 0.2 mm diam. Acorns solitary or clustered, subsessile; cup deeply cupshaped or turbinate, to 5-13 mm deep × 12-20 mm wide, scales whitish or cream, strongly tuberculate especially near base of cup; nut dark-brown, fusiform or cylyndric, 20-30 × 10-30 mm. Cotyledons distinct.


Phenology: Flowering early spring.
Habitat: Open chaparral, pinyon and juniper woodlands, desert margins, often on loose granitic soils
Elevation: 1000-1800 m

Distribution

V3 264-distribution-map.gif

Calif., Mexico (Baja California)

Discussion

Quercus cornelius-mulleri is easily distinguished from other California scrub oaks by the strongly bicolored leaves, dense minute tomentum of the abaxial leaf surface, and large acute acorns in deep tuberculate cups. Of greater difficulty are swarms of putative hybrids with Q. engelmannii, sometimes referred to as Q. ×acutidens.

In Joshua Tree National Monument a lone tree and several shrubs appear to be hybrids and backcrosses between Quercus cornelius-mulleri and Q. lobata. This tree is the basis of Quercus ×munzii J. M. Tucker.

Selected References

None.

Lower Taxa

None.
... more about "Quercus cornelius-mulleri"
acute +  and rounded +
Kevin C. Nixon +
Nixon & K. P. Steele +
attenuate-rounded;cuneate +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
2-6-lobed +
pistillate +, capitate +  and spicate +
Muller oak +
starchy +  and fleshy +
turbinate +  and cup-shaped +
×12-20 +, 5mm +  and 13mm +
multibracteate +, spiny +  and scaly +
Calif. +  and Mexico (Baja California) +
1000-1800 m +
pistillate +  and staminate +
biennial +, annual +  and maturation +
1-seeded +  and winged +
in groups +  and enclosed +
yellowish green +, grayish +  and light green +
Open chaparral, pinyon and juniper woodlands, desert margins, often on loose granitic soils +
appressed-stellate +  and stellate +
0mm;0.2mm +
minute +
pistillate +, spicate +  and staminate +
tomentulose +  and glabrate +
1.5 cm15 mm <br />0.015 m <br /> (3.5 cm35 mm <br />0.035 m <br />) +
ovate +  and oblong or narrowly obovate +
thick;10mm;20mm +
arranged +  and alternate +
irregularly +  and entire +
dark-brown +
2 cm20 mm <br />0.02 m <br /> (3 cm30 mm <br />0.03 m <br />) +
cylyndric +  and fusiform +
1 cm10 mm <br />0.01 m <br /> (3 cm30 mm <br />0.03 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
Flowering early spring. +
cream +  and whitish +
connate +  and distinct +
few-to-many +
tuberculate +
reduced +
anastomosing +, branching +  and unbranched +
Illustrated +
inconspicuous +
distinct +
subulate +  and dilated +
enlarged +
0 cm0 mm <br />0 m <br /> (0.02 cm0.2 mm <br />2.0e-4 m <br />) +
Quercus cornelius-mulleri +
Quercus sect. Quercus +
species +
spheric +  and ovoid terete or angled +
bristle-tipped +
mucronate +
brownish +, yellowish +  and gray +
0.1 cm1 mm <br />0.001 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
glabrate +  and tomentulose +
200 cm2,000 mm <br />2 m <br /> (300 cm3,000 mm <br />3 m <br />) +
subevergreen +  and evergreen +