Salix brachycarpa var. psammophila

Raup

J. Arnold Arbor. 17: 230, plate 191. 1936.

Common names: Small-fruit sand dune willow
Treatment appears in FNA Volume 7. Treatment on page 87. Mentioned on page 88, 116, 122, 148.

Stems: branches densely villous; branchlets very densely woolly, villous, or long-silky. Leaves: petiole 0.8–4 mm, (densely villous to long-silky adaxially); largest medial blade oblong, elliptic to ovate, 15–34 × 7.5–19 mm, 1.4–2.6 times as long as wide, base subcordate, cordate, or convex, apex rounded or convex, abaxial surface very densely woolly, villous, or long-silky, adaxial moderately densely villous or long-silky. Catkins: staminate 9.5–24 × 6–10 mm, flowering branchlet 2–24 (–43 in later-flowering plants) mm; pistillate stout or subglobose, 18–28.5 × 7–14 mm, flowering branchlet 4–8 mm; floral bract tawny or greenish, 1.2–2.4 mm. Staminate flowers: abaxial nectary (0–) 0.9–1.5 mm, adaxial nectary 0.5–2 mm; filaments distinct, hairy throughout or on proximal 1/2; anthers (yellow), ellipsoid or shortly cylindrical, 0.5–0.8 mm. Pistillate flowers: abaxial nectary 0.3–0.6 mm, adaxial nectary narrowly oblong or ovate (with slender tips), 0.8–1.8 mm; stipe 0–0.2 mm; ovules 8–10 per ovary; styles 0.4–1.2 mm. Capsules 4.5–6.5 mm.


Phenology: Flowering late Jun-early Aug.
Habitat: Open sand dunes
Elevation: 200-300 m

Discussion

Of conservation concern.

Previously, I (G. W. Argus 1965) did not recognize var. psammophila, but field experience showed that this sand dune endemic deserves formal taxonomic status. It is mainly characterized by very dense hairiness and slightly broader leaf blades. Salix brachycarpa var. psammophila is found only on Lake Athabasca sand dunes.

Hybrids:

Variety psammophila forms natural hybrids with Salix pyrifolia, S. silicicola, and S. turnorii. All of these taxa occur together in the sand dunes at Lake Athabasca in northwestern Saskatchewan.

Variety psammophila × Salix pyrifolia resembles S. pyrifolia in its reddish petioles and midribs, margins somewhat toothed, styles 0.4 mm, and stipes 0.4–1.1 mm; and S. brachycarpa var. psammophila in its sparsely hairy branches, branchlets, and ovaries.

Variety psammophila × Salix turnorii (S. ×brachypurpurea B. Boivin) resembles var. psammophila in floral bracts tawny, petioles relatively very short, and branchlets and juvenile leaves very densely long-silky; and S. turnorii in leaves amphistomatous, leaf margins serrulate, and branches yellow-brown. It is intermediate in ovaries varying from moderately densely hairy to glabrescent. The hybrids appear to be infertile (G. W. Argus 1965).

Selected References

None.

Lower Taxa

None.

"-0.48mm" is not declared as a valid unit of measurement for this property.

0 cm0 mm <br />0 m <br /> (0.09 cm0.9 mm <br />9.0e-4 m <br />) +
connate +  and distinct +
0.03 cm0.3 mm <br />3.0e-4 m <br /> (0.06 cm0.6 mm <br />6.0e-4 m <br />) +
long-silky +, villous +  and woolly +
ovate +  and oblong +
0.08 cm0.8 mm <br />8.0e-4 m <br /> (0.18 cm1.8 mm <br />0.0018 m <br />) +
long-silky;villous +
red turning yellow +
globose +, cylindrical +  and ellipsoid +
0.03 cm0.3 mm <br />3.0e-4 m <br /> (0.08 cm0.8 mm <br />8.0e-4 m <br />) +
Convex (?) +  and Rounded (?) +
George W. Argus +
convex +, cordate +  and subcordate +
Salix tribe Glaucae +
tapering +
toothed +  and 2-fid +
persistent +
redbrown +  and gray-brown +
2.4 cm24 mm <br />0.024 m <br /> (4.3 cm43 mm <br />0.043 m <br />) +
long-silky +, villous +  and woolly +
subglobose +  and globose +
0.4 cm4 mm <br />0.004 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
0.45 cm4.5 mm <br />0.0045 m <br /> (0.65 cm6.5 mm <br />0.0065 m <br />) +
pistillate +  and staminate +
1.8 cm18 mm <br />0.018 m <br /> (2.85 cm28.5 mm <br />0.0285 m <br />) +
pendulous +, spreading +  and erect +
subglobose +
0.7 cm7 mm <br />0.007 m <br /> (1.4 cm14 mm <br />0.014 m <br />) +
Small-fruit sand dune willow +
200-300 m +
on proximal 1/2 +  and hairy +
greenish +  and tawny +
0.12 cm1.2 mm <br />0.0012 m <br /> (0.24 cm2.4 mm <br />0.0024 m <br />) +
pistillate +  and staminate +
bisexual +  and unisexual +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.08 cm0.8 mm <br />8.0e-4 m <br />) +
spherical +
Open sand dunes +
straight +
unbranched +  and spicate +
subterminal +  and axillary +
membranaceous +
1.5 cm15 mm <br />0.015 m <br /> (3.4 cm34 mm <br />0.034 m <br />) +
elliptic +  and ovate +
0.75 cm7.5 mm <br />0.0075 m <br /> (1.9 cm19 mm <br />0.019 m <br />) +
arranged +  and alternate +
0.3 cm3 mm <br />0.003 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
marcescent +
deciduous +
0.1 cm1 mm <br />0.001 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
connate +  and distinct +
2-7[-10]-carpellate +
woolly +, tomentose +  and villous +
pyriform +
glandular-dotted +
flat;convex;flat;convex +
0.08 cm0.8 mm <br />8.0e-4 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
Flowering late Jun-early Aug. +
J. Arnold Arbor. +
overlapping +
distinct +
Illustrated +  and Endemic +
branched +, sympodial +, monopodial +  and branching +
decumbent;erect +
non-papillate +
cylindrical +
0 cm0 mm <br />0 m <br /> (0.02 cm0.2 mm <br />2.0e-4 m <br />) +
caducous +  and persistent +
connate +  and distinct +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.12 cm1.2 mm <br />0.0012 m <br />) +
persistent +  and deciduous +
ferruginous +  and white +
Salix brachycarpa var. psammophila +
Salix brachycarpa +
variety +
rounded +
sympodial +, branching +  and heterophyllous +
plant +  and not clonal +