Salix columbiana

(Dorn) Argus

Harvard Pap. Bot. 12: 359. 2007.

Common names: Columbia River willow
Basionyms: Salix exigua var. columbiana Dorn Brittonia 50: 204. 1998
Treatment appears in FNA Volume 7. Treatment on page 59. Mentioned on page 52, 53, 56, 58, 60.

Shrubs, 2–6.5 m. Stems: branches redbrown, glabrous or hairy at nodes; branchlets yellowbrown or redbrown, pubescent, puberulent, or glabrous, (hairs appressed, straight or wavy). Leaves: stipules usually absent or rudimentary, sometimes foliaceous on early ones (apex acute), rudimentary or foliaceous on late ones; petiole 2–5 mm, short-silky adaxially; largest medial blade (sometimes hypostomatous, glands marginal), linear or very narrowly elliptic, 58–115 × 5–17 mm, 5.7–12.8 times as long as wide, base acute or cuneate, margins flat to slightly revolute, remotely spinulose-serrulate or sinuate, apex acuminate, abaxial surface glaucous (sometimes obscured by hairs), sparsely to very densely short or long-silky, hairs appressed to slightly spreading, straight, adaxial dull, pubescent or long-silky; proximal blade margins entire or remotely denticulate; juvenile blade color obscured by hairs, densely short-silky abaxially. Catkins: staminate 20–83 × 5–13 mm, flowering branchlet 6–125 (–200) mm; pistillate densely to moderately densely flowered, slender or stout, 35–90 × 5–12 mm, flowering branchlet 11–160 mm; floral bract 1.4–4 mm, apex acute, rounded, retuse, or truncate, entire or toothed, abaxially hairy throughout or proximally, hairs straight or wavy. Staminate flowers: abaxial nectary 0.3–0.9 mm, adaxial nectary narrowly oblong, oblong, ovate, or flask-shaped, 0.5–1 mm, nectaries distinct or connate and shallowly cupshaped; filaments hairy on proximal 1/2, throughout, or basally; anthers 0.8–1.3 mm. Pistillate flowers: (abaxial nectary 0.3–0.4 mm), adaxial nectary narrowly oblong, ovate, or flask-shaped, 0.4–1.3 mm, longer than stipe, nectaries distinct or connate and shallowly or partially cupshaped; stipe 0.2–0.7 mm; ovary pyriform, long-silky, beak abruptly tapering to styles; ovules 18–30 per ovary; styles 0.1–0.2 (–0.4) mm; stigmas flat, abaxially non-papillate with rounded tip, or broadly cylindrical, (0.3–) 0.5–1.1 mm. Capsules 3.4–5.6 mm. 2n = 38.


Phenology: Flowering May-late Jul.
Habitat: Riparian, sandy-silty to sandy gravel floodplains, old beach dunes, rocky fill along streams
Elevation: 5-40 m

Discussion

Hybrids:

Salix columbiana forms natural hybrids with S. exigua var. exigua and S. sessilifolia. Both hybrids are reported from Oregon and Washington (R. D. Dorn 1998).

Selected References

None.

Lower Taxa

None.

... more about "Salix columbiana"
0.03 cm0.3 mm <br />3.0e-4 m <br /> (0.09 cm0.9 mm <br />9.0e-4 m <br />) +
long-silky +  and glaucous +
flask--shaped +, ovate +  and oblong +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.13 cm1.3 mm <br />0.0013 m <br />) +
long-silky;pubescent +
reddish turning yellow +  and yellow +
0.08 cm0.8 mm <br />8.0e-4 m <br /> (0.13 cm1.3 mm <br />0.0013 m <br />) +
Toothed (?) +, Entire (?) +, Retuse (?) +, Rounded (?) +, Acute (?) +  and Acuminate (?) +
George W. Argus +
(Dorn) Argus +
cuneate +  and acute +
Salix exigua var. columbiana +
tapering +
denticulate +  and entire +
short-silky +
toothed +  and 2-fid +
deciduous +
hairy +  and glabrous +
12.5 cm125 mm <br />0.125 m <br /> (20 cm200 mm <br />0.2 m <br />) +
glabrous +, puberulent +  and pubescent +
stout +  and slender +
1.1 cm11 mm <br />0.011 m <br /> (16 cm160 mm <br />0.16 m <br />) +
0.34 cm3.4 mm <br />0.0034 m <br /> (0.56 cm5.6 mm <br />0.0056 m <br />) +
pistillate +  and densely moderately densely flowered +
3.5 cm35 mm <br />0.035 m <br /> (9 cm90 mm <br />0.09 m <br />) +
pendulous +, spreading +  and erect +
stout +  and slender +
0.5 cm5 mm <br />0.005 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
Columbia River willow +
Oreg. +  and Wash. +
5-40 m +
0.14 cm1.4 mm <br />0.0014 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
pistillate +  and staminate +
bisexual +  and unisexual +
Riparian, sandy-silty to sandy gravel floodplains, old beach dunes, rocky fill along streams +
straight +
appressed +  and slightly spreading +
unbranched +  and spicate +
subterminal +  and axillary +
membranaceous +
hypostomatous +  and amphistomatous +
oblanceolate +, elliptic +, lorate +  and linear +
arranged +  and alternate +
marcescent +
deciduous +
58mm;115mm +
0.5 cm5 mm <br />0.005 m <br /> (1.7 cm17 mm <br />0.017 m <br />) +
flat +  and slightly revolute remotely spinulose-serrulate or sinuate +
connate +  and distinct +
2-7[-10]-carpellate +
pyriform +
glandular-dotted +
flat;convex +
0.2 cm2 mm <br />0.002 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
Flowering May-late Jul. +
Harvard Pap. Bot. +
overlapping +
distinct +
branched +, sympodial +, monopodial +  and branching +
0.03 cm0.3 mm <br />3.0e-4 m <br /> (0.05 cm0.5 mm <br />5.0e-4 m <br />) +
non-papillate +
cylindrical +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.11 cm1.1 mm <br />0.0011 m <br />) +
0.02 cm0.2 mm <br />2.0e-4 m <br /> (0.07 cm0.7 mm <br />7.0e-4 m <br />) +
caducous +  and persistent +
0.02 cm0.2 mm <br />2.0e-4 m <br /> (0.04 cm0.4 mm <br />4.0e-4 m <br />) +
distinct +  and connate +
0.01 cm0.1 mm <br />1.0e-4 m <br /> (0.02 cm0.2 mm <br />2.0e-4 m <br />) +
persistent +  and deciduous +
Salix columbiana +
Salix subg. Longifoliae +
species +
rounded +
sympodial +, branching +  and heterophyllous +