Heuchera americana var. hirsuticaulis

(Wheelock) Rosendahl

Minnesota Stud. Pl. Sci. 2: 60. 1936,.

Endemic
Basionym: Heuchera hispida var. hirsuticaulis Wheelock Bull. Torrey Bot. Club 17: 199. 1890
Synonyms: Heuchera americana var. interior Rosendahl, Butters & Lakela Heuchera hirsuticaulis
Treatment appears in FNA Volume 8. Treatment on page 95. Mentioned on page 94, 96.
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Petioles densely long-stipitate-glandular. Flowers: hypanthium free 1.1–2 mm, campanulate; petals greenish, white, or pink, narrower than sepals, margins nearly entire or finely dentate. 2n = 14.


Phenology: Flowering Apr–Jun.
Habitat: Rich woods or open grasslands often over base-saturated granite and gneiss, or in shallow rocky soil
Elevation: 200-400 m

Distribution

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Ark., Ill., Ind., Mich., Mo., Okla.

Discussion

Variety hirsuticaulis occurs in a band from Oklahoma to Michigan, where var. americana and Heuchera richardsonii overlap; it is intermediate between those taxa in floral characters but is densely long stipitate-glandular like H. richardsonii. It is probably the result of introgression from H. richardsonii from as long ago as the Pleistocene migrations. Data from a breeding study involving the two species showed a tendency for the shorter hypanthium of var. americana to dominate in the artificial hybrids between H. americana and H. richardsonii (E. F. Wells 1979).

Selected References

None.

Lower Taxa

None.

"full" is not a number."narrower" is not a number.

epigynous +, perigynous +  and hypogynous +
rounded +, obtuse +  and acute +
Elizabeth Fortson Wells +  and Barbara Greene Shipes +
- Wheelock Rosendahl +
cordate +  and nearly truncate +
Heuchera hispida var. hirsuticaulis +
not papillose +
dentate +, serrate +  and entire +
glandular-ciliate +  and ciliate +
3-9-lobed +, polygonal +, oblong +, cordate +, ovate +, orbiculate +  and reniform +
0.4 cm4 mm <br />0.004 m <br /> - 1.05 cm10.5 mm <br />0.0105 m <br /> +
3-carpellate +  and 2-carpellate +
ebracteate +, bracteate +  and 2-300 - -1000+-flowered +
Ark. +, Ill. +, Ind. +, Mich. +, Mo. +  and Okla. +
200-400 m +
unisexual +  and bisexual +
40 cm400 mm <br />0.4 m <br /> - 145 cm1,450 mm <br />1.45 m <br /> +
2-3 - -4-beaked +  and folliclelike +
Rich woods or open grasslands often over base-saturated granite and gneiss, or in shallow rocky soil +
multicellular +
short-stipitate-glandular +
campanulate +
0.11 cm1.1 mm <br />0.0011 m <br /> - 0.2 cm2 mm <br />0.002 m <br /> +
bracteate +, not secund +  and 100-1000-flowered +
dense +  and diffuse +
opposite +  and alternate +
5-9-lobed +, broadly ovate +  and cordate +
3.5 cm35 mm <br />0.035 m <br /> - 11 cm110 mm <br />0.11 m <br /> +
ovate;rounded +
dentate +  and entire +
0.01 cm0.1 mm <br />1.0e-4 m <br /> - 0.7 cm7 mm <br />0.007 m <br /> +
crassinucellate +, unitegmic +  and bitegmic +
epigynous +, perigynous +  and hypogynous +
pink +, white +  and greenish +
distinct +
unlobed +  and spatulate +
0.09 cm0.9 mm <br />9.0e-4 m <br /> - 0.4 cm4 mm <br />0.004 m <br /> +
peltate +  and jointed +
long-stipitate-glandular +
Flowering Apr–Jun. +
unequal +
Minnesota Stud. Pl. Sci. +
dark-brown +
straight +
muricate +, cellular-rugulose +, tuberculate +, pitted +, papillate +  and wrinkled +
ellipsoid +
0.06 cm0.6 mm <br />6.0e-4 m <br /> - 0.09 cm0.9 mm <br />9.0e-4 m <br /> +
green-tipped +
distinct +
0.1 cm1 mm <br />0.001 m <br /> - 0.24 cm2.4 mm <br />0.0024 m <br /> +
0.3 cm3 mm <br />0.003 m <br /> - 0.5 cm5 mm <br />0.005 m <br /> +
unbranched +  and branched +
persistent +
horizontal +  and erect +
0 cm0 mm <br />0 m <br /> - 0.01 cm0.1 mm <br />1.0e-4 m <br /> +
connate +  and distinct +
0.4 cm4 mm <br />0.004 m <br /> - 0.7 cm7 mm <br />0.007 m <br /> +
short-stipitate-glandular +  and glabrous +
Heuchera americana var. interior +  and Heuchera hirsuticaulis +
Heuchera americana var. hirsuticaulis +
Heuchera americana +
variety +
glandular-ciliate +  and ciliate +
crenate +  and dentate +
annual +, biennial +  and perennial +