Cyperus odoratus

Linnaeus

Sp. Pl. 1: 46. 1753.

Common names: Rusty flatsedge
Illustrated
Synonyms: Cyperus acicularis (Nees) Steudel Cyperus californicus S. Watson Cyperus eggersii Boeckeler Cyperus engelmannii Steudel Cyperus ferax Richard Cyperus ferax subsp. engelmannii (Steudel) Kükenthal Cyperus ferax subsp. speciosus (Vahl) Kükenthal Cyperus ferruginescens (Kunth) M. C. Johnston Cyperus haenkei Liebmann Cyperus huarmensis Schultes Cyperus macrocephalus Britton Cyperus michauxianus Vahl Cyperus odoratus var. engelmannii (Steudel) J. Rich. Carter, S. D. Jones & Wipff Cyperus oxycarioides Schrader ex Nees Cyperus speciosus (Linnaeus) Schrader ex Nees Cyperus squarrosus var. parvus Britton Diclidium aciculare Kunth Diclidium odoratum (Boeckeler) C. B. Clarke Mariscus ferax (Richard) Urban Mariscus huarmensis (Liebmann) T. Koyama Torulinium eggersii (Schultes) C. B. Clarke Torulinium ferax (Linnaeus) S. S. Hooper Torulinium macrocephalum Torulinium michauxianum Torulinium odoratum
Treatment appears in FNA Volume 23. Treatment on page 191. Mentioned on page 145, 146, 184, 188.
Revision as of 00:39, 30 July 2020 by imported>Volume Importer
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Herbs, annual (or short-lived perennial). Culms trigonous, (4–) 10–50 (–130) cm × (0.5–) 1–4 mm. Leaves flanged V or inversely W-shaped, 5–30 (–60) cm × 4–12 mm. Inflorescences: spikes 1–5 (–12), loosely to densely cylindric, ovoid, or ± pyramidal, 10–20 (–40) × (8–) 10–30 (–35) mm; rays (0–2) 6–9 (–12), 2–8 (–13) cm; if rays absent, infloresence a single dense, capitate cluster of closely imbricate spikes; bracts (4–) 5–8 (–10), horizontal to ascending at 30–60°, (3–) 10–25 (–55) cm × 1–14 mm; rachilla hyaline or thickened bronze, carmine, or yellowish, wings clasping achene. Spikelets (10–) 20–60 (–100), linearoblong to narrowly linear, cylindric to slightly flattened (when scale tips spreading), (5–) 8–15 (–38) × 0.8–1.3 (–1.9) mm; floral scales (4–) 8–12– (30), medially green, laterally reddish to stramineous to shiny brown or beige, medially 2–5-ribbed, laterally 1–3-ribbed, ovate to elliptic, (2–) 2.2–2.8 (–3.2) × (1.2–) 1.4–1.6 (–1.8) mm, apex entire or emarginate with mucro to 0.3 mm. Flowers: anthers (0.2–) 0.3–0.4 (–0.7) mm; styles 0.4–0.7 (–1) mm; stigmas (1–) 1.5–3 mm. Achenes brown, reddish-brown, or black, stipitate, narrowly ellipsoid to oblong (rarely obovoid-oblong), (1–) 1.2–1.5 (–1.9) × 0.5–0.6 (–0.75) mm, stipe 0.1–0.2 mm, apex acute to barely obtuse, surfaces finely papillose.


Phenology: Fruiting summer–early fall.
Habitat: Emergent shorelines, disturbed, muddy places, fresh or slightly brackish marshes
Elevation: 0–1500 m

Distribution

V23 331-distribution-map.jpg

Ont., Que., Ala., Ariz., Ark., Calif., Colo., Conn., Del., D.C., Fla., Ga., Ill., Ind., Iowa, Kans., Ky., La., Maine, Md., Mass., Mich., Minn., Miss., Mo., Nebr., N.H., N.J., N.Mex., N.Y., N.C., Ohio, Okla., Oreg., Pa., R.I., S.C., S.Dak., Tenn., Tex., Vt., Va., Wash., W.Va., Wis., Central America, South America, Asia, Africa, Pacific Islands (Hawaii), Australia

Discussion

As circumscribed here, Cyperus odoratus is easily identified by its cylindric to subcylindric spikelets in which the corky rachilla of the mature spikelet disarticulates at the base of each scale. The mature spikelet breaks into segments each consisting of a scale and an internode of the rachilla clasping the achene with its corky wings. Cyperus odoratus is an exceedingly variable pantropical and warm-temperate species. Numerous segregates have been named, some of which may deserve recognition when the species is studied in detail worldwide.

Selected References

None.

Lower Taxa

None.

"shortened" is not a number.

... more about "Cyperus odoratus"
0.15 cm1.5 mm <br />0.0015 m <br /> (0.19 cm1.9 mm <br />0.0019 m <br />) +
0.06 cm0.6 mm <br />6.0e-4 m <br /> (0.075 cm0.75 mm <br />7.5e-4 m <br />) +
black +, reddish-brown +  and brown +
0.12 cm1.2 mm <br />0.0012 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
narrowly ellipsoid;oblong +
0.5mm;0.6mm +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.07 cm0.7 mm <br />7.0e-4 m <br />) +
0.03 cm0.3 mm <br />3.0e-4 m <br /> (0.04 cm0.4 mm <br />4.0e-4 m <br />) +
emarginate +  and entire +
acute;barely obtuse +
Gordon C. Tucker* +, Brian G. Marcks* +  and J. Richard Carter * +
Linnaeus +
persistent +  and deciduous +
decurrent +
Undefined subg. Diclidium +
w--shaped +  and keeled +
glumaceous +  and foliaceous +
2-ranked +  and arranged +
25 cm250 mm <br />0.25 m <br /> (55 cm550 mm <br />0.55 m <br />) +
10 cm100 mm <br />0.1 m <br /> (25 cm250 mm <br />0.25 m <br />) +
horizontal +  and ascending +
0.1 cm1 mm <br />0.001 m <br /> (1.4 cm14 mm <br />0.014 m <br />) +
parallel +  and divergent +
terete +, rolled +  and plicate +
Rusty flatsedge +
50 cm500 mm <br />0.5 m <br /> (130 cm1,300 mm <br />1.3 m <br />) +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
10 cm100 mm <br />0.1 m <br /> (50 cm500 mm <br />0.5 m <br />) +
scabridulous +
trigonous +
1mm;4mm +
Ont. +, Que. +, Ala. +, Ariz. +, Ark. +, Calif. +, Colo. +, Conn. +, Del. +, D.C. +, Fla. +, Ga. +, Ill. +, Ind. +, Iowa +, Kans. +, Ky. +, La. +, Maine +, Md. +, Mass. +, Mich. +, Minn. +, Miss. +, Mo. +, Nebr. +, N.H. +, N.J. +, N.Mex. +, N.Y. +, N.C. +, Ohio +, Okla. +, Oreg. +, Pa. +, R.I. +, S.C. +, S.Dak. +, Tenn. +, Tex. +, Vt. +, Va. +, Wash. +, W.Va. +, Wis. +, Central America +, South America +, Asia +, Africa +, Pacific Islands (Hawaii) +  and Australia +
0–1500 m +
0.28 cm2.8 mm <br />0.0028 m <br /> (0.32 cm3.2 mm <br />0.0032 m <br />) +
0.16 cm1.6 mm <br />0.0016 m <br /> (0.18 cm1.8 mm <br />0.0018 m <br />) +
laterally reddish +  and stramineous +
0.22 cm2.2 mm <br />0.0022 m <br /> (0.28 cm2.8 mm <br />0.0028 m <br />) +
ovate +  and elliptic +
0.14 cm1.4 mm <br />0.0014 m <br /> (0.16 cm1.6 mm <br />0.0016 m <br />) +
hypogynous +  and subtending +
biconvex +  and trigonous +
Emergent shorelines, disturbed, muddy places, fresh or slightly brackish marshes +
subumbellate +  and capitate +
spreading +  and erect +
multi-ranked +, 2-ranked +, 3-ranked +  and alternate +
30 cm300 mm <br />0.3 m <br /> (60 cm600 mm <br />0.6 m <br />) +
5 cm50 mm <br />0.05 m <br /> (?) +  and 30 cm300 mm <br />0.3 m <br /> (?) +
w--shaped +  and flanged +
0.4 cm4 mm <br />0.004 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
0 cm0 mm <br />0 m <br /> (0.03 cm0.3 mm <br />3.0e-4 m <br />) +
with (1-)3-6(-30) bristles and/or scales +
Fruiting summer–early fall. +
2-3(-4)-carpellate +
antrorse +  and extrorse +
3 (?) +  and 1 (?) +
yellowish +, carmine +, bronze +, thickened +  and hyaline +
8 cm80 mm <br />0.08 m <br /> (13 cm130 mm <br />0.13 m <br />) +
unequal +
single +, 6 +  and 9 +
2 cm20 mm <br />0.02 m <br /> (8 cm80 mm <br />0.08 m <br />) +
adventitious +
basal +  and proximal +
cylindric;compressed +
cylindric +
W2 +, W1 +  and Illustrated +
2 cm20 mm <br />0.02 m <br /> (4 cm40 mm <br />0.04 m <br />) +
3 cm30 mm <br />0.03 m <br /> (3.5 cm35 mm <br />0.035 m <br />) +
1 cm10 mm <br />0.01 m <br /> (2 cm20 mm <br />0.02 m <br />) +
pyramidal;ovoid;pyramidal;ovoid;cylindric +
1 cm10 mm <br />0.01 m <br /> (3 cm30 mm <br />0.03 m <br />) +
1.5 cm15 mm <br />0.015 m <br /> (3.8 cm38 mm <br />0.038 m <br />) +
0.13 cm1.3 mm <br />0.0013 m <br /> (0.19 cm1.9 mm <br />0.0019 m <br />) +
0.8 cm8 mm <br />0.008 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
20 +  and 60 +
cylindric;slightly flattened +
0.8mm;1.3mm +
septate +, hollow +  and solid +
compressed +, terete +  and trigonous +
0.1 cm1 mm <br />0.001 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
papillate +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
0.01 cm0.1 mm <br />1.0e-4 m <br /> (?) +  and 0.02 cm0.2 mm <br />2.0e-4 m <br /> (?) +
0.07 cm0.7 mm <br />7.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
2-3-fid +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.07 cm0.7 mm <br />7.0e-4 m <br />) +
papillose +
Cyperus acicularis +, Cyperus californicus +, Cyperus eggersii +, Cyperus engelmannii +, Cyperus ferax +, Cyperus ferax subsp. engelmannii +, Cyperus ferax subsp. speciosus +, Cyperus ferruginescens +, Cyperus haenkei +, Cyperus huarmensis +, Cyperus macrocephalus +, Cyperus michauxianus +, Cyperus odoratus var. engelmannii +, Cyperus oxycarioides +, Cyperus speciosus +, Cyperus squarrosus var. parvus +, Diclidium aciculare +, Diclidium odoratum +, Mariscus ferax +, Mariscus huarmensis +, Torulinium eggersii +, Torulinium ferax +, Torulinium macrocephalum +, Torulinium michauxianum +  and Torulinium odoratum +
Cyperus odoratus +
Cyperus subg. Diclidium +
species +
tuberous +, stoloniferous +  and rhizomatous +