Clematis subg. Viorna
Syn. Fl. N. Amer. 1(1): 5. 1895.
Woody vines or erect, ± herbaceous perennials, clumped (or patch-forming from rhizomes in C. socialis). Leaves simple. Leaf-blade 1-pinnate or 2-pinnate, ternate, or finely dissected; ultimate divisions lobed or unlobed, margins entire or few-toothed. Inflorescences terminal and/or axillary, on current-years stems; 3-7-flowered bracteate cymes or flowers solitary or paired, peduncles bracteate [or several-many-flowered panicles]. Flowers bisexual, usually nodding (± erect in some spp., esp. C. morefieldii); sepals ascending, connivent at least proximally and usually much of length, variously colored, lanceolate or oblong to broadly ovate, thick, usually leathery, abaxially glabrous to silky, hirsute, or tomentose; filaments slender, usually pubescent (except C. pitcheri var. dictyota), connectives often ± prolonged (especially in C. pitcheri); staminodes absent. Achenes flattened; beak variable in length, plumose to nearly glabrous.
Distribution
Temperate, mostly North America, a few in Eurasia
Discussion
Species ca. 25 (18 in the flora).
Clematis integrifolia Linnaeus, with relatively wide-spreading, blue sepals, is locally naturalized in Ontario.
Selected References
None.
Lower Taxa
Key
1 | Leaves all simple, blade rarely so deeply lobed that proximal 2 lobes appear as distinct linear leaflets; plants herbaceous or ± woody at base, erect, not viny. | > 2 |
1 | At least some distal leaves of main stems distinctly compound or deeply much-dissected; plants erect or viny. | > 8 |
2 | Sepals glabrous or sparsely villous, tips acuminate; achene body cobwebby-tomentose distally, or long-pubescent. | > 3 |
2 | Sepals (except margins) minutely puberulent, silky, woolly, or nearly glabrous, tips obtuse to acute; achene body pilose or short-pilose. | > 4 |
3 | Leaf blade 3.5-11 cm wide, prominently reticulate adaxially; beak not plumose, proximally silky-tomentose, sparsely appressed-pubescent to nearly glabrous distally; Kans., Nebr., Mo. | Clematis fremontii |
3 | Leaf blade 0.2-2(-3.5) cm wide, not prominently reticulate adaxially; beak distinctly plumose; Fla. | Clematis baldwinii |
4 | Leaf blade abaxially moderately to densely soft-pubescent, rarely nearly glabrous; secondary and tertiary veins forming prominent reticulum on adaxial surface. | > 5 |
4 | Leaf blade abaxially glabrous or nearly so, sparsely or rarely densely villous on veins; secondary and tertiary veins not forming prominent reticulum on adaxial surface. | > 6 |
5 | Stems and abaxial surface of leaf blades moderately silky-pilose with spreading hairs, rarely nearly glabrous; beak yellowish brown to reddish brown; hairs of achene rim appressed-ascending. | Clematis ochroleuca |
5 | Stems and abaxial surface of leaf blades densely silky-tomentose with appressed hairs; beak white to pale yellow; hairs of achene rim spreading. | Clematis coactilis |
6 | Sepals abaxially silky to woolly; beak white to pale yellow. | Clematis albicoma |
6 | Sepals abaxially glabrous to minutely puberulent; beak coppery brown. | > 7 |
7 | Leaf blade abaxially glaucous and glabrous; sepals abaxially glabrous. | Clematis addisonii |
7 | Leaf blade abaxially not glaucous, sparsely to densely villous on veins; sepals abaxially nearly glabrous or minutely puberulent. | Clematis viticaulis |
8 | Plants erect or sprawling, not viny; stems to 0.65m. | > 9 |
8 | Plants viny, petioles and/or rachises of leaves often functioning as tendrils; stems usually 1-5m. | > 13 |
9 | Larger leaf blades (1-)2-3-pinnate or -ternate or deeply dissected. | > 10 |
9 | Leaf blade 1-pinnate or simple. | > 11 |
10 | Leaflets usually less than 1.5 cm wide, mostly more than 2.5 times as long as wide, mostly unlobed, if lobed then with lateral lobes 1 or 2, usually small, distinctly narrower than central portion; blade abaxially sparsely to densely hirsute; beak plumose. | Clematis hirsutissima |
10 | Leaflets usually more than 1.5 cm wide and/or less than 2.5 times as long as wide, mostly lobed, lateral lobes often nearly as wide as central portion; blade glabrous or nearly so (rachis and petiolules may be ±hirsute); beak glabrous or inconspicuously appressed-pubescent. | Clematis bigelovii |
11 | Terminal leaflets usually tendril-like; blades of simple leaves and lateral leaflets of compound leaves usually more than 2 cm wide; sepals purple or reddish purple, whitish toward tips. | Clematis addisonii |
11 | Terminal leaflets with expanded blade, not tendril-like; blades of simple leaves and lateral leaflets usually less than 2 cm wide; sepals uniformly violet-blue. | > 12 |
12 | Plants strongly rhizomatous, forming patches; sepals 2-2.5(-3) cm; beak 1.5-2.5 cm. | Clematis socialis |
12 | Plants not rhizomatous; sepals 2.5-5.5 cm; beak 6-10 cm. | Clematis baldwinii |
13 | Largest leaf blades 1-2-pinnate, leaflets mostly deeply lobed; beak 1-3 cm, inconspicuously appressed-pubescent to nearly glabrous. | Clematis pitcheri |
13 | Largest leaves simple, blades 1-pinnate, or if some 2-pinnate, leaflets of 2-pinnate leaves usually unlobed, rarely 2-5-lobed; beak 2-7 cm, plumose (appressed-puberulent in C.crispa and C.pitcheri). | > 14 |
14 | Leaflets abaxially glaucous and glabrous, rarely with a few scattered hairs. | > 15 |
14 | Leaflets not glaucous, usually abaxially pubescent, sometimes glabrous. | > 18 |
15 | Leaves mostly simple, only distalmost compound. | Clematis addisonii |
15 | Leaves all or mostly compound, simple leaves, if any, only on small branches and near base of main stem. | > 16 |
16 | Leaf blade ±thin, secondary and tertiary veins not forming prominent reticulum on adaxial surface. | Clematis glaucophylla |
16 | Leaf blade leathery, secondary and tertiary veins forming prominent reticulum on adaxial surface. | > 17 |
17 | Sepals rose-red to scarlet abaxially and at tip adaxially; tips recurved. | Clematis texensis |
17 | Sepals pale lavender or blue-violet to reddish purple abaxially, often paler or greenish adaxially, tips slightly spreading. | Clematis versicolor |
18 | Sepals with thin, crispate margins to 6 mm wide distally. | Clematis crispa |
18 | Sepals without expanded, thin, crispate margins or with margins less than 2.5 mm wide. | > 19 |
19 | Leaf blade leathery, secondary and tertiary veins forming prominent reticulum on adaxial surface. | > 20 |
19 | Leaf blade thin, secondary and tertiary veins not forming prominent reticulum on adaxial surface. | > 21 |
20 | Beak plumose, with spreading hairs; leaf blade finely reticulate (ultimate closed areoles mostly less than 2 mm in longer dimension), even quaternary veins distinctly raised on adaxial surface. | Clematis reticulata |
20 | Beak sparsely pubescent to silky, with ascending to appressed hairs; leaf blade less finely reticulate (ultimate closed areoles mostly more than 2 mm in longer dimension), tertiary and quaternary veins scarcely or not raised on adaxial surface. | Clematis pitcheri |
21 | Stems generally cobwebby as well as villous; bracts near base of peduncle/pedicel; sepals densely silky-pubescent abaxially, pinkish suffused with green, tips spreading to short-reflexed. | Clematis morefieldii |
21 | Stems without cobwebby pubescence; bracts well above base of peduncle/pedicel; sepals sparsely pubescent abaxially, pale lavender to reddish purple, tips recurved. | Clematis viorna |
"connivent" is not a number."thick" is not a number.