Taxon	Illustrator ⠉
	Caulophyllum thalictroides Nandina domestica	John Myers John Myers

Herbs, perennial, deciduous, to 2-9 dm, glabrous. Rhizomes nodose, producing 2 leaves per year. Aerial stems present. Leaves caducous, cauline, 2-ranked, 1st leaf larger than 2d leaf, 2-4-ternately compound; petioles short or absent. Leaf-blade broadly obovate in overall outline; rachis pulvinate; leaflet blades broadly obovate, entire or lobed, margins not toothed; venation pinnate to palmate. Inflorescences terminal, compound cymes. Flowers 3-merous, 6-20 mm; bracteoles caducous, 3-4, sepaloid; sepals 6, yellow, purple, red, brown, or green, petaloid; petals 6, fan-shaped, bearing nectar; stamens 6; anthers dehiscing by 2 apically hinged flaps; pollen exine reticulate; ovary bladderlike; placentation appearing basal; styles eccentric. Fruits not developed, ovary wall soon rupturing. Seeds 2, elevated on elongating stalk, naked at maturity; seed-coat blue, fleshy, glaucous; aril absent. x = 8.

Distribution

North America, Asia

Discussion

Species 3 (2 in the flora).

Caulophyllum species are understory herbs of mesophytic forests, alluvial flats, rich mesic slopes, and limestone slopes. The seeds of Caulophyllum are buoyant and showy and may be dispersed by water as well as other means; seed germination is hypogeal, the cotyledons remaining underground. Caulophyllum is occasionally cultivated in woodland gardens.

Selected References

None.

Lower Taxa

Caulophyllum giganteum, Caulophyllum thalictroides

Key

1	Pistil 3–5 mm; style 1–2 mm; stamen filaments 1.5–2.5 mm; sepals 6–9 mm; inflorescences with 4–18 flowers; 1st leaf (2–)3-ternate; leaflets 5–10 cm.	Caulophyllum giganteum
1	Pistil 1–3 mm; style 0.25–1 mm; stamen filaments 0.5–1.5 mm; sepals 3–6 mm; inflorescences with 5– 70 flowers; 1st leaf 3(–4)-ternate; leaflets 3–8 cm.	Caulophyllum thalictroides