Eriogonum lonchophyllum

Torrey & A. Gray

Proc. Amer. Acad. Arts 8: 173. 1870.

Endemic
Synonyms: Eriogonum ainsliei Wooton & Standley Eriogonum corymbosum var. humivagans (Reveal) S. L. Welsh Eriogonum effusum subsp. ainsliei (Wooton & Standley) S. Stokes Eriogonum effusum subsp. fendlerianum (Bentham) S. Stokes Eriogonum effusum var. nudicaule Torrey Eriogonum effusum subsp. salicinum (Greene) S. Stokes Eriogonum fendlerianum (Torrey) Small Eriogonum humivagans Greene Eriogonum intermontanum Reveal Eriogonum lonchophyllum var. fendlerianum (Bentham) Reveal Eriogonum lonchophyllum var. humivagans (Reveal) Reveal Eriogonum lonchophyllum var. intermontanum (Reveal) Reveal Eriogonum lonchophyllum var. nudicaule (Torrey) Reveal Eriogonum lonchophyllum var. saurinum (Reveal) S. L. Welsh Eriogonum nudicaule Eriogonum nudicaule subsp. scoparium (Small) S. Stokes Eriogonum nudicaule subsp. tristichum (Small) S. Stokes Eriogonum salicinum Eriogonum saurinum Eriogonum scoparium Eriogonum tristichum
Treatment appears in FNA Volume 5. Treatment on page 262. Mentioned on page 232, 236, 237, 239, 240, 256, 257, 263, 273.
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Shrubs, subshrubs, or herbs, spreading to erect, not scapose, (1–) 1.5–5 × 2–5 (–8) dm, glabrous or rarely floccose to tomentose, grayish. Stems spreading or erect, usually without persistent leaf-bases, up to 1/2 or more height of plant; caudex stems absent or matted to spreading; aerial flowering-stems spreading to erect, slender, solid, not fistulose, 0.3–3 dm, glabrous or, rarely, floccose to tomentose, tomentose among leaves. Leaves basal or cauline on proximal 1/2 of stem, 1 per node; petiole 0.5–2 cm, tomentose to floccose or glabrous; blade narrowly lanceolate or oblanceolate to elliptic, 1.5–7 (–9) × 0.2–2 cm, velvety to densely white-tomentose abaxially, sparsely tomentose to thinly floccose or glabrous and green adaxially, margins plane or occasionally crenulate. Inflorescences cymose, dense to more commonly open, 2–25 × 2–20 cm; branches dichotomous, glabrous or, rarely, floccose; bracts 3, scalelike, usually triangular, and 1–3 mm, or occasionally leaflike, 8–30 mm, and otherwise similar to leaf-blades. Peduncles absent or erect, 0.1–0.8 cm, glabrous. Involucres 1 per node or 2–5 per cluster, turbinate to turbinate-campanulate, 2.5–4 × (1.3–) 1.5–3.5 (–4) mm, glabrous; teeth 5, erect, 0.4–0.9 mm. Flowers 2–3.5 (–4) mm; perianth white, glabrous; tepals connate proximal 1/4–1/3, monomorphic, oblanceolate, elliptic to oblong or obovate; stamens exserted, 2–4 mm; filaments pilose proximally. Achenes light-brown to brown, 2–3 mm, glabrous except (typically) for slightly papillate beak.


Phenology: Flowering Jun–Oct.
Habitat: Heavy gumbo clay soil or (at higher elevations) sandy-loam to gravelly or rocky soil and outcrops, mixed grassland, saltbush, blackbrush, and sagebrush communities, pinyon-juniper and montane conifer woodlands
Elevation: 1400-2900 m

Discussion

Eriogonum lonchophyllum is widespread and occasionally rather common in sagebrush-dominated communities in the Rocky Mountains of Colorado, northern New Mexico and eastern Utah. The distribution is fragmented, many of the populations are markedly distinct, and several have been provided with names. The continued recognition of E. coloradense is probably dubious given what is already reduced here to synonymy under E. lonchophyllum. Eriogonum lonchophyllum typically has glabrous flowering stems and inflorescence branches, yet floccose to tomentose individuals are known from Eagle (Reveal & Davidse 861, UTC) and Garfield (Goodrich & D. Nelson 24582, BRY) counties in Colorado. Plants representing the typical expression of the species differ from the majority of populations assigned here in having broad leaves and a sprawling habit. Such plants grow in deep shade under tall conifers and are known currently only from the type locality. Variation in the size, shape, and distribution of leaves has been used to distinguish varieties of E. lonchophyllum. At their extremes, the differences are remarkable. For example, plants with linear, basal leaves have been named var. nudicaule. Such plants are found in the New Mexico counties of Rio Arriba, Sandoval, and Santa Fe. Another herbaceous phase also with basal, but broader leaves and known from the Book Cliffs and Tavaputs Plateau regions of northeastern Emery, southern Uintah, and northern Grand counties, Utah, and in Rio Blanco County, Colorado, has been named var. intermontanum. The low, spreading subshrubby phase found along the foothills of the Front Ranges in south-central Colorado (El Paso, Fremont, Jefferson, Las Animas, Otero, and Pueblo counties) and adjacent north-central New Mexico (Colfax and Taos counties) is known as var. fendlerianum. These plants have velvety, tomentose leaf blades. Another shrubby phase is found in and around the Dinosaur National Park in Daggett and Uintah counties, Utah, and in northwestern Rio Blanco County, Colorado. This is var. saurinum, characterized by having sheathing leaf blades along the flowering stems that vary from narrowly oblanceolate to narrowly elliptic; it is occasionally confused with E. corymbosum in that area. At the north end of Mesa Verde National Park and on Sleeping Ute Mountain of Montezuma County, Colorado, is a large shrub that outwardly resembles E. corymbosum in size and shape. It is similar to, but larger than, the more eastern var. fendlerianum. The most common phase of the species (as represented by var. humivagans, including E. salicinum, E. scoparium, and E. tristichum) is found mainly on fine, clayey to gravelly, alluvial soils in the Rocky Mountains and western desert ranges of Colorado (Archuleta, Delta, Dolores, Eagle, Garfield, Gunnison, La Plata, Mesa, Moffat, Montezuma, Montrose, Ouray, Rio Blanco, Saguache, and San Miguel counties), northern New Mexico (Rio Arriba, Sandoval, and San Juan counties), and San Juan County, Utah. The majority of the plants have long, narrow leaf blades (2–5 mm wide), but populations with broader leaves are common in southwestern Colorado. It is this latter expression of E. lonchophyllum that can be difficult to distinguish from E. corymbosum in portions of eastern Utah and western Colorado. A single collection (Rammel s.n., 1872, MIN, US) was supposedly gathered in Texas. Until this is confirmed, the species is not considered to be part of the Texas flora.

Selected References

None.

Lower Taxa

None.

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... more about "Eriogonum lonchophyllum"
light-brown;brown +
included +  and exserted +
3-gonous +
usually red +  and cream or yellow +
oblong +  and ellipsoid or oval +
acute;obtuse or rounded +
James L. Reveal +
Torrey & A. Gray +
deciduous +  and persistent +
stipelike +
Undefined subg. Eucycla +
papillate +
awn-tipped +  and simple +
7 cm70 mm <br />0.07 m <br /> (9 cm90 mm <br />0.09 m <br />) +
1.5 cm15 mm <br />0.015 m <br /> (7 cm70 mm <br />0.07 m <br />) +
velvety +  and densely white-tomentose abaxially +
oblanceolate;elliptic +
0.2 cm2 mm <br />0.002 m <br /> (2 cm20 mm <br />0.02 m <br />) +
not awn-tipped +
glandular +, pubescent +  and glabrous +
triangular +  and scale-like +
0.8 cm8 mm <br />0.008 m <br /> (3 cm30 mm <br />0.03 m <br />) +
not brittle +
spreading +  and erect +
floccose +, , +  and glabrous +
scabrellous +
ridged +, angled +  and round +
whorled +, opposite +  and alternate +
Colo. +, N.Mex. +  and Utah. +
1400-2900 m +
0.35 cm3.5 mm <br />0.0035 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
unisexual +, , +  and bisexual +
not attenuate +
0.2 cm2 mm <br />0.002 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
decumbent to erect +  and prostrate +
not fistulose +
unwinged +  and winged +
black +, red +, brown +  and yellowish +
exserted +  and included +
pubescent +  and glabrous +
spheroidal +, 4-gonous +, lenticular +, biconvex +, discoid +, 3-gonous +  and 2-gonous +
Heavy gumbo clay soil or (at higher elevations) sandy-loam to gravelly or rocky soil and outcrops, mixed grassland, saltbush, blackbrush, and sagebrush communities, pinyon-juniper and montane conifer woodlands +
reduced +
open +  and cymose +
capitate +  and subcapitate +
2 cm20 mm <br />0.02 m <br /> (25 cm250 mm <br />0.25 m <br />) +
cymelike +, paniclelike +, racemelike +  and spikelike +
2 cm20 mm <br />0.02 m <br /> (20 cm200 mm <br />0.2 m <br />) +
connate +  and free +
0.35 cm3.5 mm <br />0.0035 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
turbinate +  and turbinate-campanulate +
nonmembranous +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
in whorls +, opposite +, alternate +  and rosulate +
persistent +
cauline +  and basal +
bractlike +
pubescent +  and glabrous +
crenulate +  and plane +
not swollen +
orthotropous +
accrescent +
curved +  and straight +
erect to deflexed +
0.1 cm1 mm <br />0.001 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
cylindric +  and urceolate +
accrescent +
fleshy +, indurate +  and membranous +
with extrafloral nectaries +  and not articulate +
tomentose +  and floccose or glabrous +
0.5 cm5 mm <br />0.005 m <br /> (2 cm20 mm <br />0.02 m <br />) +
Flowering Jun–Oct. +
homostylous +  and 3-carpellate +
Proc. Amer. Acad. Arts +
, +  and solid +
slender;stout +
persistent +
0.2 cm2 mm <br />0.002 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
decumbent to erect +  and prostrate +
erect +  and spreading +
glandular +, pubescent +  and glabrous +
penicillate +, fimbriate +  and capitate +
deciduous +  and persistent +
2-lobed +  and cylindric +
coriaceous +  and membranous +
consisting of a series +  and tubular +
distinct +
erect;spreading or recurved +
spreading;erect +
Eriogonum ainsliei +, Eriogonum corymbosum var. humivagans +, Eriogonum effusum subsp. ainsliei +, Eriogonum effusum subsp. fendlerianum +, Eriogonum effusum var. nudicaule +, Eriogonum effusum subsp. salicinum +, Eriogonum fendlerianum +, Eriogonum humivagans +, Eriogonum intermontanum +, Eriogonum lonchophyllum var. fendlerianum +, Eriogonum lonchophyllum var. humivagans +, Eriogonum lonchophyllum var. intermontanum +, Eriogonum lonchophyllum var. nudicaule +, Eriogonum lonchophyllum var. saurinum +, Eriogonum nudicaule +, Eriogonum nudicaule subsp. scoparium +, Eriogonum nudicaule subsp. tristichum +, Eriogonum salicinum +, Eriogonum saurinum +, Eriogonum scoparium +  and Eriogonum tristichum +
chambered +  and solid +
slender +  and stout +
Eriogonum lonchophyllum +
Eriogonum subg. Eucycla +
species +
emarginate +  and entire +
0 +  and 1/2 +
1/4 +  and 1/3 +
elliptic +  and oblong or obovate +
coriaceous +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.09 cm0.9 mm <br />9.0e-4 m <br />) +
persistent +
membranous +
pubescent +  and glabrous +
plant +, herb +, subshrub +  and shrub +
rarely floccose +  and tomentose +
synoecious +, monocarpic +, polycarpic +  and , +
3 +, 2 +  and 1 +