Cyperus papyrus

Linnaeus

Sp. Pl. 1: 47. 1753.

Common names: Papyrus Egyptian paper-reed
IntroducedIllustrated
Treatment appears in FNA Volume 23. Treatment on page 173. Mentioned on page 147, 167, 174.

Herbs, perennial, rhizomatous, stout. Culms roundly trigonous, 300–500 cm × 15–45 mm, glabrous. Leaves bladeless. Inflorescences: spikes loosely cylindric, 10–20 × 6–10 mm; rays 40–100, drooping or arching, slender, 10–30 cm; 2d order rays 8–20 cm; bracts 4–10, ± erect, V-shaped, 3–8 cm × 4–15 mm; 2d order bracts 2–5, (1.5–) 4–16 cm × 0.5–2 mm; rachilla persistent, separating laterally, remaining firmly attached basally, wings 0.3–0.4 mm wide. Spikelets 6–30, slightly compressed, linear, ± quadrangular, 6–10 × 0.8–1 mm; floral scales 6–16, reddish beside 5-ribbed green medial part, white to hyaline near margins, ovate-elliptic, 1.8–2.2 × 1.2–1.5 mm, apex acute to obtuse. Flowers: anthers 0.8–1 mm (connectives prolonged beyond anther as red subulate appendage 0.2–0.5 mm, apex usually setose); styles 0.2–0.4 mm; stigmas 0.8–1.2 mm. Achenes pale-brown, sessile, oblong, 0.8–1 × 0.4 mm, apex scarcely apiculate, surfaces puncticulate.


Phenology: Fruiting summer.
Habitat: Stream banks, marshes
Elevation: 0–30 m

Distribution

V23 287-distribution-map.jpg

Introduced; Fla., s Europe, sw Asia, Africa

Discussion

Cyperus papyrus is conspicuous in the field by its great height, leafless culm, and open, hemispheric inflorescence with drooping rays.

Ancient Egyptians, Greeks, and Romans prepared paper from the pith of the culms of Cyperus papyrus, which is abundant along the Nile River.

Selected References

None.

Lower Taxa

None.

"shortened" is not a number.

... more about "Cyperus papyrus"
0.08 cm0.8 mm <br />8.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
0.08 cm0.8 mm <br />8.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
acute;obtuse +
Gordon C. Tucker* +, Brian G. Marcks* +  and J. Richard Carter * +
Linnaeus +
persistent +  and deciduous +
decurrent +
w--shaped +  and keeled +
glumaceous +  and foliaceous +
2-ranked +  and arranged +
1.5 cm15 mm <br />0.015 m <br /> (4 cm40 mm <br />0.04 m <br />) +
4 cm40 mm <br />0.04 m <br /> (16 cm160 mm <br />0.16 m <br />) +
v--shaped +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
parallel +  and divergent +
terete +, rolled +  and plicate +
Papyrus +  and Egyptian paper-reed +
300 cm3,000 mm <br />3 m <br /> (500 cm5,000 mm <br />5 m <br />) +
scabridulous +
trigonous +
15mm;45mm +
Fla. +, s Europe +, sw Asia +  and Africa +
0–30 m +
deciduous +  and persistent +
hypogynous +  and subtending +
biconvex +  and trigonous +
Stream banks, marshes +
subumbellate +  and capitate +
spreading +  and erect +
multi-ranked +, 2-ranked +, 3-ranked +  and alternate +
white +  and hyaline +
0.18 cm1.8 mm <br />0.0018 m <br /> (0.22 cm2.2 mm <br />0.0022 m <br />) +
ovate-elliptic +
0.12 cm1.2 mm <br />0.0012 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
with (1-)3-6(-30) bristles and/or scales +
Fruiting summer. +
2-3(-4)-carpellate +
antrorse +  and extrorse +
3 (?) +  and 1 (?) +
continuous +, articulate +, winged +  and wingless +
0.3mm +  and 0.4mm +
conspicuous +
unequal +
arching +  and drooping +
40 +  and 100 +
slender +
8 cm80 mm <br />0.08 m <br /> (20 cm200 mm <br />0.2 m <br />) +
adventitious +
basal +  and proximal +
cylindric;compressed +
cylindric +
Introduced +  and Illustrated +
1 cm10 mm <br />0.01 m <br /> (2 cm20 mm <br />0.02 m <br />) +
cylindric +
0.6 cm6 mm <br />0.006 m <br /> (1 cm10 mm <br />0.01 m <br />) +
0.6 cm6 mm <br />0.006 m <br /> (1 cm10 mm <br />0.01 m <br />) +
6 +  and 30 +
quadrangular;compressed +
0.8mm;1mm +
septate +, hollow +  and solid +
compressed +, terete +  and trigonous +
papillate +
0.08 cm0.8 mm <br />8.0e-4 m <br /> (0.12 cm1.2 mm <br />0.0012 m <br />) +
2-3-fid +
0.02 cm0.2 mm <br />2.0e-4 m <br /> (0.04 cm0.4 mm <br />4.0e-4 m <br />) +
Undefined subg. Chlorocyperus +, Cyperus subg. Mariscus +  and Undefined subg. Papyrus +
Cyperus papyrus +
Cyperus subg. Cyperus +
species +