Eriogonum exaltatum

M. E. Jones

Contr. W. Bot. 15: 61. 1929.

Common names: Ladder wild buckwheat
Endemic
Treatment appears in FNA Volume 5. Treatment on page 397. Mentioned on page 384, 398.

Herbs, erect, annual, (0.5–) 3–6 (–10) dm, glabrous, glaucous, grayish. Stems: caudex absent; aerial flowering-stems erect, solid, not fistulose, (0.5–) 1–2 dm, glabrous. Leaves basal; petiole 1–10 cm, tomentose; blade subcordate to orbiculate, (1.5–) 2–5 (–8) × (1.5–) 2–5 (–8) cm, densely white-tomentose abaxially, floccose to subglabrous and green adaxially, margins often wavy. Inflorescences cymose, spreading when immature, quickly becoming narrowly erect and strict with whiplike branches, (5–) 10–50 (–80) × 10–50 cm; branches glabrous, glaucous; bracts 3, scalelike, 1–1.5 (–2) × 0.4–1 mm. Peduncles absent or erect, straight, slender, 0.1–0.2 cm, glabrous. Involucres turbinate, 2–2.5 (–3) × 1.5–2.5 mm, glabrous; teeth 5, erect, 0.5–1.5 mm. Flowers 1.5–2 mm; perianth white with green or reddish midribs, becoming pinkish, glabrous; tepals dimorphic, those of outer whorl oblong, those of inner whorl lanceolate; stamens exserted, 1.5–2.5 mm; filaments pilose proximally. Achenes dark-brown to blackish, 3-gonous, 2–2.5 mm, glabrous. 2n = 40.


Phenology: Flowering May–Oct.
Habitat: Sandy to gravelly flats and slopes, saltbush, creosote bush, greasewood, blackbrush, and mesquite communities, rarely in pinyon-juniper woodlands
Elevation: 500-1400 m

Distribution

V5 818-distribution-map.gif

Ariz., Calif., Nev., Utah.

Discussion

Eriogonum insigne has been a troublesome taxon.  The type, collected by E. Palmer in 1876, is a curious specimen from near Paragonah in Iron County, Utah.  That expression has not been recollected and, after another frustrating review of the type material, it is now referred to E. deflexum, along with other odd specimens from southern Nevada and California that have always been difficult to assign.  As a result, the tall, upright plants with long, whiplike branches of northwestern Arizona (Mohave County), southern Nevada (northeastern Clark and southern Lincoln counties), and southwestern Utah (southwestern Washington County), are now recognized under the name E. exaltatum.  Some specimens remain problematic.  Immature plants of E. exaltatum (Goodding 2302, GH, MIN, MO, NY, RM, UC) resemble E. bifurcatum. Specimens from Eureka Valley (DeDecker 4741, NY) and western Pahrump Valley (Abrams 14248, DS, GH, NY) in Inyo County, California, appear to belong to E. exaltatum.

Like Eriogonum exaltatum, some populations of E. deflexum var. deflexum have nearly erect involucres.  The plants from the Buried Hills (Nye and Clark counties, Nevada) are particularly odd.  Single plants from a few collections in Kane County, Utah, have sessile, seemingly erect involucres in the forks of inflorescence branches.  None of these plants has the characteristic long, whiplike branches of E. exaltatum, and they are not included in E. deflexum var. deflexum. The type of E. deflexum var. rectum from San Bernardino County, California (Reveal & Broome 6385, CAS, NY, US, etc.), has individual specimens with both deflexed and somewhat erect involucres, along with whiplike branches.  Collections from Imperial and San Diego counties, previously associated with what is here termed E. exaltatum, are now considered specimens of E. deflexum.  Clearly, more work is required to understand these species fully.

Selected References

None.

Lower Taxa

None.

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... more about "Eriogonum exaltatum"
dark-brown;blackish +
included +  and exserted +
3-gonous +
usually red +  and cream or yellow +
oblong +  and ellipsoid or oval +
James L. Reveal +
M. E. Jones +
deciduous +  and persistent +
stipelike +
Eriogonum sect. Ganysma +
awn-tipped +  and simple +
5 cm50 mm <br />0.05 m <br /> (8 cm80 mm <br />0.08 m <br />) +
5 cm50 mm <br />0.05 m <br /> (8 cm80 mm <br />0.08 m <br />) +
2 cm20 mm <br />0.02 m <br /> (5 cm50 mm <br />0.05 m <br />) +
floccose +  and subglabrous +
subcordate;orbiculate +
2 cm20 mm <br />0.02 m <br /> (5 cm50 mm <br />0.05 m <br />) +
not awn-tipped +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
0.1 cm1 mm <br />0.001 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
glandular +, pubescent +  and glabrous +
scale-like +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
grooved +, disarticulating +  and dichotomous +
50 cm500 mm <br />0.5 m <br /> (80 cm800 mm <br />0.8 m <br />) +
not brittle +
10 cm100 mm <br />0.1 m <br /> (50 cm500 mm <br />0.5 m <br />) +
glaucous +  and glabrous +
scabrellous +
whip-like +
10 cm100 mm <br />0.1 m <br /> (50 cm500 mm <br />0.5 m <br />) +
spreading +  and erect +
whorled +, opposite +  and alternate +
Ladder wild buckwheat +
Ariz. +, Calif. +, Nev. +  and Utah. +
500-1400 m +
unisexual +, , +  and bisexual +
not attenuate +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
decumbent to erect +  and prostrate +
not fistulose +
slender;stout +
unwinged +  and winged +
black +, red +, brown +  and yellowish +
exserted +  and included +
pubescent +  and glabrous +
spheroidal +, 4-gonous +, lenticular +, biconvex +, discoid +, 3-gonous +  and 2-gonous +
Sandy to gravelly flats and slopes, saltbush, creosote bush, greasewood, blackbrush, and mesquite communities, rarely in pinyon-juniper woodlands +
capitate +  and subcapitate +
racemose +, paniculate +  and cymose-paniculate +
erect +  and spreading +
cymelike +, paniclelike +, racemelike +  and spikelike +
connate +  and free +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
nonmembranous +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
in whorls +, opposite +, alternate +  and rosulate +
marcescent +
persistent +
bractlike +
pubescent +  and glabrous +
reddish +  and green +
not swollen +
orthotropous +
accrescent +
erect to deflexed +
filiform;capillary +
0.1 cm1 mm <br />0.001 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
pinkish;white +
cylindric +  and urceolate +
accrescent +
fleshy +, indurate +  and membranous +
with extrafloral nectaries +  and not articulate +
1 cm10 mm <br />0.01 m <br /> (10 cm100 mm <br />0.1 m <br />) +
Flowering May–Oct. +
homostylous +  and 3-carpellate +
Contr. W. Bot. +
, +  and solid +
slender;stout +
persistent +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
decumbent to erect +  and prostrate +
spreading +  and erect +
glabrous +, short-hispid +, villous +, hirsute +, floccose +  and tomentose +
penicillate +, fimbriate +  and capitate +
deciduous +  and persistent +
2-lobed +  and cylindric +
coriaceous +  and membranous +
consisting of a series +  and tubular +
distinct +
erect;spreading or recurved +
spreading;erect +
chambered +  and solid +
slender +  and stout +
Eriogonum exaltatum +
Eriogonum subg. Ganysma +
species +
emarginate +  and entire +
1/5 +  and 1/2 +
lobed +  and laciniate +
coriaceous +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
persistent +
membranous +
pubescent +  and glabrous +
annual +  and perennial +
glaucous +  and glabrous +
synoecious +, monocarpic +, polycarpic +  and , +
3 +, 2 +  and 1 +