Acaena pinnatifida

Ruiz & Pavón

Fl. Peruv. 1: 68, plate 104, fig. b. 1798. 1798

Illustrated
Synonyms: Acaena californica Bitter A. pinnatifida var. californica (Bitter) Jepson
Treatment appears in FNA Volume 9. Treatment on page 324.

Plants from branched woody caudex. Stems decumbent, becoming suberect, 1.5–5 dm (in fruit), 4 mm diam.; branches erect or leafy. Leaves (crowded at bases of stems); stipules absent; petiole to 4 cm; blade 3–15 cm, leaflets 6–10 per side, gray-green (at least on dried specimens), ovate, 4–10 mm, pinnatisect, lobes 3–7 (–11), linear, margins revolute, apex penicillate (having a little brush or tuft of hairs at tips), abaxial surface sericeous, adaxial sparsely pilose. Scapes terminal, to 50 cm, 1.5–3 mm diam., sparsely pilose, proximal scape and leaf-sheaths with multicellular hairs among 1-celled hairs, with (0–) 2–3 cauline leaves to 10 cm. Inflorescences 15–35-flowered, interrupted spikes, (flowers crowded distally, 2–15 cm in flower, elongating to 50 cm in fruit); bracteoles leaflike to linear, ciliate. Flowers 6 mm diam.; sepals 4–5, elliptic, 3 mm, abaxially pilose, adaxially glabrous; stamens usually 4, filaments 2–3 mm, anthers dark red; stigma rose, laciniate. Fruiting hypanthia ovoid, 3–4 mm diam., surfaces tomentose, obscurely 4-angled, bearing 10–20 spines in longitudinal ranks, to 3 [–4] mm, spines with retrorse hairs or barbs.


Phenology: Flowering Mar–Jun.
Habitat: Coastal grasslands, open rocky slopes, sand dunes, old roadsides
Elevation: 50–400 m

Distribution

V9 529-distribution-map.jpg

Calif., South America (Argentina), South America (Chile)

Discussion

Herbarium records indicate occurrences of Acaena pinnatifida between Santa Barbara and Sonoma counties.

Selected References

None.

Lower Taxa

None.

"dm" is not declared as a valid unit of measurement for this property."dm" is not declared as a valid unit of measurement for this property.

... more about "Acaena pinnatifida"
Bryony Macmillan +
Ruiz & Pavón +
compound +  and simple +
opposite +  and alternate +
15 cm150 mm <br />0.15 m <br /> (18 cm180 mm <br />0.18 m <br />) +
spatulate +
3 cm30 mm <br />0.03 m <br /> (15 cm150 mm <br />0.15 m <br />) +
coriaceous +
leaflike +  and linear +
free +  and distinct +
Calif. +, South America (Argentina) +  and South America (Chile) +
not +  and aggregated +
50–400 m +
0.2 cm2 mm <br />0.002 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
6 cm60 mm <br />0.06 m <br /> (?) +
unisexual +  and bisexual +
not +  and aggregated +
spindle--shaped +
Coastal grasslands, open rocky slopes, sand dunes, old roadsides +
1-celled +  and multicellular +
globose +
gray +  and redbrown +
0.3 cm3 mm <br />0.003 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
glabrous +  and tomentose +
Angled (?) +, Obconic (?) +, Globose (?) +, Oblong (?) +, Ovoid (?) +  and Obtriangular (?) +
axillary +  and terminal +
crassinucellate +
persistent +
0.4 cm4 mm <br />0.004 m <br /> (1 cm10 mm <br />0.01 m <br />) +
entire +, dentate +  and pinnatisect +
incised +, revolute +  and flat +
inferior +  and superior +
clustered +, biseriate +  and superposed +
0 (?) +  and 4 (?) +
free +  and distinct +
5 +  and 4 +
0 cm0 mm <br />0 m <br /> (4 cm40 mm <br />0.04 m <br />) +
Flowering Mar–Jun. +
adnate +, free +, connate +  and distinct +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
0 cm0 mm <br />0 m <br /> (50 cm500 mm <br />0.5 m <br />) +
not arillate +
persistent +
free +  and distinct +
glabrous +  and pilose +
triangular +  and elliptic +
3 cm30 mm <br />0.03 m <br /> (?) +
Illustrated +
glandular +  and eglandular +
glabrous +  and pubescent +
free +  and distinct +
4 cm40 mm <br />0.04 m <br /> (?) +
1 +  and 20 +
laciniate +
red +  and white +
deciduous +
distinct +
basal +, lateral +, subterminal +  and terminal +
fimbriate +
0.3 cm3 mm <br />0.003 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
0 cm0 mm <br />0 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
Acaena californica +  and A. pinnatifida var. californica +
Acaena pinnatifida +
species +
inconspicuous +
enlarged +  and small +
0.3 cm3 mm <br />0.003 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +