Boerhavia coulteri var. coulteri

Illustrated
Synonyms: Boerhavia rosei Standley
Treatment appears in FNA Volume 4. Treatment on page 27.

Stems 2–8 dm. Leaf-blades 10–50 × 6–32 mm. Flowers: pedicel 0.2–1.6 mm; bracts at base of perianth usually 2; perianth 1–2 mm distal to constriction; stamens 2–3 (–4), slightly exserted. Inflorescences terminal clusters, axis 15–60 mm. Fruits (4–) 7–20 (–22) per cluster, usually overlapping by 50–100% of their length, or 2–4 in group, separated by small gap from next group and with distal fruits overlapping, pale tan, narrowly obpyramidal, 2.5–3.2 (–3.6) × (0.9–) 1.1 (–1.4) mm (l/w: [2.2–] 2.5–3.1 [–3.3]), apex bluntly conic-truncate, round-truncate, or truncate; ribs obtuse, each often with sharp ridge, slightly rugose near sulci; sulci 0.1–0.2 times as wide as base of ribs, slightly rugose.


Phenology: Flowering late spring–early fall.
Habitat: Sandy or loamy soils in arid grasslands, among brush, roadsides in deserts and cultivated areas
Elevation: [100-]500-1400 m

Distribution

V4 42-distribution-map.gif

Ariz., N.Mex., Utah, Mexico (Sinaloa), Mexico (Sonora)

Discussion

Most plants of Boerhavia coulteri var. coulteri have fruit apices that are bluntly conic- or round-truncate. Along the Santa Cruz River particularly, some plants have abruptly truncate fruits; those plants represent the phase known as B. rosei. Early in the twentieth century, J. J. Thornber proposed the name “B. standleyi” for such plants but never published it. P. C. Standley (1911) described the ribs of B. rosei as narrow, adding in 1918 that the sulci were broad and open. As the type demonstrates, the relative breadth of the ribs and sulci is exactly the reverse, confusing regional botanists for several generations.

Selected References

None.

Lower Taxa

None.

"thin" is not a number."dm" is not declared as a valid unit of measurement for this property.

with few coarse hairs +, puberulent +  and glabrous +
hirtellous +  and glabrous +
truncate +, round-truncate +  and conic-truncate +
Richard W. Spellenberg +
(Hooker f.) S. Watson +
1.5 cm15 mm <br />0.015 m <br /> (6 cm60 mm <br />0.06 m <br />) +
glutinous +
symmetric +  and asymmetric +
deciduous +
round +, obtuse +  and acute +
unequal +
woody +, leathery +  and fleshy +
Senkenbergia coulteri +
1 cm10 mm <br />0.01 m <br /> (5 cm50 mm <br />0.05 m <br />) +
deltate;deltate-ovate;deltate;deltate-ovate;oval;ovate;lanceolate +
6mm;32mm +
deciduous +  and persistent +
distinct +
ovate +, linear-lanceolate +, lance-acuminate +  and lanceolate +
minute +  and not accrescent +
corollalike +  and petaloid +
0.32 cm3.2 mm <br />0.0032 m <br /> (0.36 cm3.6 mm <br />0.0036 m <br />) +
0.11 cm1.1 mm <br />0.0011 m <br /> (0.14 cm1.4 mm <br />0.0014 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (0.32 cm3.2 mm <br />0.0032 m <br />) +
0.09 cm0.9 mm <br />9.0e-4 m <br /> (0.11 cm1.1 mm <br />0.0011 m <br />) +
Ariz. +, N.Mex. +, Utah +, Mexico (Sinaloa) +  and Mexico (Sonora) +
[100-]500-1400 m +
curved +  and straight +
inconspicuous +  and showy +
1 +  and 80 +
chasmogamous +  and bisexual +
0.32 cm3.2 mm <br />0.0032 m <br /> (0.36 cm3.6 mm <br />0.0036 m <br />) +
0.11 cm1.1 mm <br />0.0011 m <br /> (0.14 cm1.4 mm <br />0.0014 m <br />) +
2.5mm;3.2mm +
2 (?) +  and 4 (?) +
obpyramidal +
coriaceous +
1.1 cm11 mm <br />0.011 m <br /> (?) +
small +
Sandy or loamy soils in arid grasslands, among brush, roadsides in deserts and cultivated areas +
thyrsiform +, cymose +  and pedunculate +
2-5 times with sticky internodal bands +
1 cm10 mm <br />0.01 m <br /> (5 cm50 mm <br />0.05 m <br />) +
0.6 cm6 mm <br />0.006 m <br /> (3.2 cm32 mm <br />0.032 m <br />) +
unequal +
(4-)5-lobed +
pubescent +  and glabrous +
crisped +  and sinuate +
0.02 cm0.2 mm <br />2.0e-4 m <br /> (?) +  and 0.16 cm1.6 mm <br />0.0016 m <br /> (?) +
campanulate +
constricted +
0.1 cm1 mm <br />0.001 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (3 cm30 mm <br />0.03 m <br />) +
Flowering late spring–early fall. +
Proc. Amer. Acad. Arts +
glandular-pubescent +, glabrous +  and smooth +
obtuse +
thick +  and thin +
fibrous;fleshy +
membranaceous +
ribbed +, 10-ribbed +  and 5-ribbed +
pubescent +, glabrous +  and warty +
wrinkled +
Illustrated +
erect +  and decumbent-ascending +
fusiform +  and linear +
filiform +
0.1-0.2 times as wide as base +
Boerhavia rosei +
woody +  and soft +
Boerhavia coulteri var. coulteri +
Boerhavia coulteri +
variety +
umbellate +, subumbellate +, spicate +  and cymose +
capitate +  and subcapitate +
singly +  and racemose +
gland-tipped +
expanded +
pubescent +, glabrous +  and glandular +
fusiform +  and ropelike +