Cercocarpus betuloides
in J. Torrey and A. Gray, Fl. N. Amer. 1: 427. 1840.
Shrubs or trees, 10–50 (–80) dm, sparsely to moderately branched, young stems, leaves, pedicels, and hypanthia sericeous (hairs straight, antrorse, appressed, or ± ascending), or hirsute-pilose (hairs erect, straight), or ± villous (hairs wavy or wavy-crinkled), or hairs tightly coiled or 1/2 coiled, or 1/2 coiled at base and erect-ascending, wavy distally, 0.2–0.8 (–1.3) mm. Stems: long-shoot internodes 3–20 (–55) mm, ± sparsely hairy, glabrescent; short-shoots 3–30 (–55) × 1–2.5 mm. Leaves persistent or ± drought-deciduous; stipules (1.5–) 3.5–6 (–10) mm, 1.5–2.5 mm in s California; petiole 1.5–9 (–16) mm; blade oblong-obovate, rhombic, suborbiculate to broadly ovate, rarely oblongelliptic, (8–) 12–50 (–78) × (4–) 8–35 (–64) mm, membranous to coriaceous, base cuneate to rounded, rarely ± cordate, margins ± revolute or flat, dentate, serrate, or, in broader leaves, crenate in distal 1/5–3/4, apex rounded, sometimes acute, abaxial surface sericeous, pilose, villous, or with curled hairs on veins, areoles canescent or glabrous, adaxial sparsely hairy, sometimes glabrate. Flowers 1–18 per short-shoot; hypanthial tubes sericeous, woolly, or villous or with coiled or spreading hairs; hypanthial cups 2–4 × 4–7 mm; sepals 5, broadly deltate, 1–2.5 mm, acute-acuminate; stamens 20–61, anthers 1–1.5 mm, hirsute. Achenes 7–11 (–13.5) × 1.5–2.5 (–2.9) mm; fruiting pedicels (1.5–) 3–11 (–21) mm; hypanthial tubes (5–) 7–13 (–17) mm; pedicel/tube ratio (15–) 35–90 (–126) %; fruit awns 5–8.5 (–12.5) cm, proximal setae 2.7–5 mm.
Distribution
Ariz., Calif., Oreg., nw Mexico
Discussion
Varieties 3 (3 in the flora).
The western, spring-flowering Cercocarpus betuloides in the broad sense, centered in California, is well supported by molecular data. A total of five taxa have been recognized in recent floras; molecular data provide no support for these taxa, but instead support recognition of a single inclusive taxon.
Selected References
None.
Lower Taxa
Key
| 1 | Leaves, young stems, and hypanthial tubes appressed-sericeous or hypanthial tubes also with erect-ascending hairs (hairs not coiled); flowers 1–3(–10) per short shoot; transmontane s, c California, c Arizona, and sw Oregon. | Cercocarpus betuloides var. betuloides |
| 1 | Leaves, young stems, and hypanthial tubes commonly with tightly coiled hairs (except in some insular plants where sericeous); flowers (1–)3–6(–18) per short shoot; cismontane s California | > 2 |
| 2 | Leaf blades sparsely to moderately hairy, subcoriaceous to coriaceous, margins serrate or dentate or crenate in distal 1/5–3/4; coastal s California and adjacent islands. | Cercocarpus betuloides var. blancheae |
| 2 | Leaf blades abaxially densely woolly, strongly coriaceous, margins (often strongly revolute, usually obscuring teeth), crenate, dentate-serrate in distal 1/2–3/4; sw Santa Catalina Island. | Cercocarpus betuloides var. traskiae |
"dm" is not declared as a valid unit of measurement for this property.