Codriophorus sect. Codriophorus
Stem sparingly or freely irregularly, dichotomously or fasciculately branched, sometimes filiform and almost unbranched, absent short, lateral tuft-like branchlets. Leaves lingulate, elliptic, broadly ovate, ovate or oblong-lanceolate to broadly or occasionally narrowly lanceolate; margins 1–2 (–4) -stratose, recurved, reflexed, incurved or plane on both sides; apices narrowly to broadly rounded-obtuse or long acuminate to acute, muticous or very rarely shortly hyaline-tipped, entire or sharply to bluntly, remotely irregularly toothed, erose-dentate, sinuate to cristate in the upper 1/5–1/4; costa strong and broad, ceasing in mid leaf to subpercurrent, in transverse-section 2–4-stratose in the distal and medial parts, weakly to strongly convex, reniform, crescent-shaped to semiterete abaxially, flat or convex on the adaxial side, in the basal part 3–6 (–7) -layered, situated at the bottom of a shallow or deeper, usually wide-angled groove, lunate, reniform or rectangular and distinctly flattened, strongly convex on the abaxial side; basal marginal laminal cells not differentiated; distal laminal cells 1-stratose to variously 2-stratose, irregularly roundedquadrate, subquadrate, oval to rounded-hexagonal, strongly papillose to smooth, straight-walled in the distal part.
Distribution
North America, South America (Colombia), Eurasia, n Africa, Atlantic Islands (Azores), Atlantic Islands (Canary Islands), Atlantic Islands (Iceland), Atlantic Islands (Madeira)
Discussion
Species 9 (6 in the flora).
Members of sect. Codriophorus are recognized by their lingulate, broadly ovate to ovate- or oblong-lanceolate leaves, isodiametric and straight-walled distal laminal cells, and costa that is broad and robust, usually over 100 µm wide at the base. The costa is situated in a shallow, wide-angled groove and is mostly 2–4-stratose in the distal and medial parts, 3–6(–7)-stratose in the basal part, and anatomically strongly differentiated.
Selected References
None.
Lower Taxa
Key
1 | Laminal cells smooth or very slightly papillose on young leaves; leaf margins entire or with a few irregular teeth at the apex. | Codriophorus depressus |
1 | Laminal cells distinctly papillose; leaf margins remotely irregularly toothed, erose-dentate, sinuate to cristate in the distal 1/5-1/4, if entire then broadly rounded-obtuse | > 2 |
2 | Leaves lanceolate; apex irregularly bluntly erose-dentate; costa markedly convex and crescent-shaped abaxially. | Codriophorus ryszardii |
2 | Leaves lingulate, elliptical to broadly ovate or ovate-lanceolate; apex broadly rounded-obtuse to acute, irregularly dentate to entire | > 3 |
3 | Costa subpercurrent, entire and only shortly forked at the tip | > 4 |
3 | Costa extending 1/2-3/4 way up the leaf, distinctly spurred | > 5 |
4 | Laminal cells strongly papillose, 1- or very rarely 2-stratose; leaves unbordered. | Codriophorus acicularis |
4 | Laminal cells indistinctly papillose over the cell walls, always variously 2-stratose in the distal part; leaves distinctly bordered with fleshy and broad limbidia. | Codriophorus norrisii |
5 | Leaf apex broadly rounded, flat to subcucullate, always entire or sometimes sinuate to remotely bluntly erose-dentate. | Codriophorus mollis |
5 | Leaf apex broadly rounded-obtuse to subacute, often tubular, strongly dentate or very rarely subentire. | Codriophorus aduncoides |