Cylindropuntia californica

(Torrey & A. Gray) F. M. Knuth in C. Backeberg and F. M. Knuth

in C. Backeberg and F. M. Knuth, Kaktus-ABC, 125. 1935.

Common names: California cholla
Illustrated
Basionym: Cereus californicus Torrey & A. Gray Fl. N. Amer. 1: 555. 1840 Opuntia californica (Torrey & A. Gray) Coville 1899
Treatment appears in FNA Volume 4. Treatment on page 113. Mentioned on page 103, 104.

Shrubs, decumbent to erect, essentially trunkless, 0.3–2.5 m. Stem segments green to purplish green, 6–30 × 1.5–2.5 cm; tubercles prominent, oval to elongate-elliptic, 0.7–3 cm, sometimes appearing as ribs; areoles elliptic, 4.5–5 × 3–3.5 mm; wool tan, aging dark gray. Spines (0–) 7–13 (–20) per areole, at most areoles, rarely obscuring stems, yellow to orangebrown with yellow tips, aging dark gray, subequal or 1–6 (–7) much longer to 3 cm; sheaths whitish or gold to brown, moderately loose. Glochids yellow-tan to redbrown in dense adaxial crescent and scattered, stouter ones at basal margins. Flowers: inner tepals yellow to greenish yellow; outer ones tipped red, 15–30 mm; filaments green or yellow; anthers yellow; style light green or cream-white (rarely pinkish) apically; stigma lobes cream-white to yellow. Fruits green to yellowish, drying gray-tan, short top-shaped to subspheric, 15–32 × 13–26 mm, leathery-dry, tuberculate, few-spined or spineless; proximal tubercles longest; umbilicus deep; rim slightly flanged; areoles 13–30. Seeds tan, squarish to angular in outline, warped, (4–) 5–7 × 4–5 mm, sides with 2–4 deep depressions; girdle smooth.

Discussion

Varieties 4 (2 in the flora).

Selected References

None.

Key

1 Shrubs, suberect to decumbent; stem tubercles oval, shorter, usually 0.7-1.5 cm; spines/areole usually subequal Cylindropuntia californica var. californ
1 Shrubs, suberect to erect; stem tubercles elliptic, longer, usually 1.5-3 cm; spines/areole commonly with 1(-3) central spines much longer Cylindropuntia californica var. parkeri
0 cm0 mm <br />0 m <br /> (3 cm30 mm <br />0.03 m <br />) +
cushionlike +
gray +  and tan +
0.07 cm0.7 mm <br />7.0e-4 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
deciduous +  and persistent +
0.45 cm4.5 mm <br />0.0045 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
13 +  and 30 +
obdeltate +  and rhombic +
0.3 cm3 mm <br />0.003 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
Donald J. Pinkava +
(Torrey & A. Gray) F. M. Knuth in C. Backeberg and F. M. Knuth +
Cereus californicus +  and Opuntia californica +
proliferating +
flattened +  and cylindric +
California cholla +
Calif. +  and Mexico +
yellow +  and green +
1.5 cm15 mm <br />0.015 m <br /> (3 cm30 mm <br />0.03 m <br />) +
15 cm150 mm <br />0.15 m <br /> (30 cm300 mm <br />0.3 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (15 cm150 mm <br />0.15 m <br />) +
spineless +  and few-spined +
gray-tan;green;yellowish +
indehiscent +
persistent +, long +  and deciduous +
15mm;32mm +
tuberculate +
top-shaped +  and subspheric +
dry +  and fleshy +
1.3 cm13 mm <br />0.013 m <br /> (2.6 cm26 mm <br />0.026 m <br />) +
deciduous +
flattened +  and conic +
0 cm0 mm <br />0 m <br /> (3 cm30 mm <br />0.03 m <br />) +
succulent +
tuberculate +
fluted;spheric;depressed-spheric or club-shaped +
tinged color +
sometimes covered by outgrowths of proximal portion of style base or of flower tube wall +  and open +
persistent +  and deciduous +
ridgelike +  and nipple--shaped +
in C. Backeberg and F. M. Knuth, Kaktus-ABC, +
3;2;2;30 +
flanged +
tuberlike +
0.4 cm4 mm <br />0.004 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
warped;squarish;angular +
0.4 cm4 mm <br />0.004 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
Illustrated +
deciduous +  and persistent +
conspicuous +
7 (?) +  and 13 (?) +
flattened +
hard +  and rigid +
decurrent +
winged +  and unsegmented +
tuberculate +  and smooth +
40 cm400 mm <br />0.4 m <br /> (50 cm500 mm <br />0.5 m <br />) +
green +  and purplish green +
6 cm60 mm <br />0.06 m <br /> (30 cm300 mm <br />0.3 m <br />) +
1.5 cm15 mm <br />0.015 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
segmented +
woody +  and succulent +
cream-white +  and yellow +
cream-white +  and light green +
Opuntia sect. Cylindropuntia +
adventitious +
tuberlike +
epipetric +  and epiphytic +
Cylindropuntia californica +
Cylindropuntia +
species +
dull +  and glossy +
oval +  and elongate-elliptic +
0.7 cm7 mm <br />0.007 m <br /> (3 cm30 mm <br />0.03 m <br />) +
terrestrial;epiphytic or epipetric +
decumbent +  and erect +
dark gray +  and tan +