Delphinium subsect. Echinata

(Ewan) N. I. Malyutin

Bot. Zhurn. (Moscow & Leningrad) 72: 689. 1987.

Basionym: Delphinium series Echinatae Ewan Bull. Torrey Bot. Club 69: 139. 1942
Treatment appears in FNA Volume 3.

Roots 3-9 (-20) -branched, 3-8 (-30) cm, fascicled, dry to fleshy, ± succulent, thin threadlike segments restricted to termini of major segment; buds minute. Stems 1 (-2) per root, unbranched, elongation delayed 2-10 weeks after leaf initiation; base usually not narrowed, firmly attached to root; proximal internodes much shorter than those of midstem. Leaves basal and cauline, largest near base of stem, others often abruptly smaller on distal portion of stem; basal petioles spreading, cauline petioles ascending; blades of basal leaves more rounded and with fewer, wider lobes than cauline leaves. Inflorescences usually with 2-11 flowers per 5 cm, dense to open, cylindric, spurs rarely intersecting rachis; pedicel ascending, usually less than 2 cm, rachis to midpedicel angle more than 30°; bracts markedly smaller and fewer lobed than leaves. Fruits erect. Seeds ± rectangular, 1.5-2.5 × 0.8-1.5 mm, not ringed at proximal end, ± wing-margined; seed-coats without wavy ridges (sometimes with prism-shaped structures in D. hansenii), cells ± brick-shaped, cell margins straight or undulate.

Discussion

Species 4 (4 in the flora).

Selected References

None.

Key

1 Proximal petioles glabrous to puberulent with short (less than 0.5 mm) appressed and/or arched hairs. > 2
1 Proximal petioles with long (more than 0.5 mm), straight spreading hairs. > 3
2 Sepal color and/or stem pubescence different from that of other individuals with which they occur; stem base puberulent with short arched hairs. Delphinium hesperium subsp. pallescens
2 Sepal color and stem pubescence similar to that of other individuals with which they occur; stem base glabrous to pubescent with short straight hairs. Delphinium hesperium
3 Seeds echinate, appearing fuzzy to naked eye; lateral sepals 13 mm or shorter; flowers usually more than 12 per main inflorescence branch. Delphinium hansenii
3 Seeds not echinate, appearing ± smooth to naked eye; lateral sepals 10 mm or longer; flowers usually fewer than 12 per main inflorescence branch. > 4
4 Stem base longitudinally ridged. Delphinium hesperium
4 Stem base not longitudinally ridged. > 5
5 Margins of lower petals ciliate; bracteoles usually 7 mm or less from sepals; sepal spur straight. Delphinium variegatum
5 Margins of lower petals glabrous; bracteoles usually 7 mm or more from sepals; sepal spur often downcurved for more than 3 mm at apex. Delphinium hutchinsoniae

"/" is not declared as a valid unit of measurement for this property."thin" is not a number.

Michael J. Warnock +
(Ewan) N. I. Malyutin +
usually larger +
narrowed +
expanded +
Delphinium series +
straight +
0.2 cm2 mm <br />0.002 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
smaller +
subopposite-subalternate +
minute +
0 cm0 mm <br />0 m <br /> (?) +  and 0.3 cm3 mm <br />0.003 m <br /> (?) +
globose;cylindric +
brick--shaped +
sessile +  and pedicellate +
showy +  and inconspicuous +
sessile +  and aggregate +
curved-cylindric +
axillary +  and terminal +
less ovate +  and elliptic +
0.8 cm8 mm <br />0.008 m <br /> (1.8 cm18 mm <br />0.018 m <br />) +
round +  and pentagonal or reniform +
whorled +, opposite +  and alternate +
cauline +  and basal +
largest +
lacerate;crenate +
clawed +, 2-lobed +, ovate +  and plane +
0.2 cm2 mm <br />0.002 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
palmate +  and pinnate +
incised;toothed;incised;toothed;entire +
smaller +
0 cm0 mm <br />0 m <br /> (2 cm20 mm <br />0.02 m <br />) +
distinct +
conspicuous +
spurred +, funnel--shaped +, cup-shaped +  and plane +
reduced +
Bot. Zhurn. (Moscow & Leningrad) +
2-100-flowered +
5 cm50 mm <br />0.05 m <br /> (40 cm400 mm <br />0.4 m <br />) +
3-9(-20)-branched +
8 cm80 mm <br />0.08 m <br /> (30 cm300 mm <br />0.3 m <br />) +
tuberlike +
3 cm30 mm <br />0.03 m <br /> (8 cm80 mm <br />0.08 m <br />) +
succulent +, dry +  and fleshy +
wing-margined +
dark-brown +  and black +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
not ringed +
rectangular +
0.08 cm0.8 mm <br />8.0e-4 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
thread-like +
yellow;red;yellow;red;white;purple;blue +
not persistent +
distinct +
3 +  and 6 +
not +  and veined +
persistent +
Delphinium subsect. Echinata +
Delphinium sect. Diedropetala +
subsection +
0.8 cm8 mm <br />0.008 m <br /> (2.4 cm24 mm <br />0.024 m <br />) +
herbaceous +  and woody +