Dodecatheon pulchellum var. macrocarpum

(A. Gray) Reveal

Sida 22: 863. 2006 ,.

Common names: Alaskan shootingstar
Basionym: Dodecatheon meadia var. macrocarpum A. Gray in W. H. Brewer et al., Bot. California 1: 467. 1876
Synonyms: Dodecatheon macrocarpum var. alaskanum Hultén Dodecatheon pauciflorum var. alaskanum (Hultén) C. L. Hitchcock Dodecatheon pulchellum subsp. alaskanum (Hultén) Hultén Dodecatheon pulchellum var. alaskanum (Hultén) B. Boivin Dodecatheon pulchellum subsp. macrocarpum (A. Gray) Roy L. Taylor & MacBryde Dodecatheon pulchellum subsp. superbum (Pennell & Stair) Hultén Primula pauciflora var. macrocarpa (A. Gray) Mast & Reveal
Treatment appears in FNA Volume 8. Treatment on page 283. Mentioned on page 280, 281.

Plants glabrous. Leaves (3–) 5–20 (–35) × (0.5–) 1.5–5 cm; blade elliptic or narrowly ovate to ovate. Pedicels glabrous. Flowers: calyx glabrous; corolla-tube yellow with red to maroon, thin, wavy ring, lobes magenta to lavender, 7–18 (–20) mm; filament tube yellow, 1.7–2.7 mm; anthers (4.5–) 5–8.5 mm; pollen-sacs usually yellow, at least apically, sometimes maroon or faintly purplish, often with pink to maroon speckles or lines abaxially, connective maroon. 2n = 88, 132.


Phenology: Flowering late spring–summer.
Habitat: Moist tidal flats, marshes, stream banks, slopes and cliffs mainly in meadows or grassy areas in or near conifer woodlands
Elevation: 0-1800 m

Distribution

V8 571-distribution-map.gif

B.C., Alaska, Calif., Oreg., Wash.

Discussion

Of conservation concern.

Variety macrocarpum is a coastal polyploid. It ranges from Kodiak Island of south-central Alaska southward (including many islands) through British Columbia and western Washington to western Oregon. H. J. Thompson (1953) improperly lectotypified var. macrocarpum on a W. H. Brewer (217, GH) collection from the Santa Susana Mountains, Ventura County, California, even though the name was based solely upon a W. H. Dall (GH) collection from Middleton Island, Valdez-Cordova County, Alaska. The coastal phase in Alaska has larger leaves and generally taller plants; away from the immediate coast, it grades quickly into the more typical, shorter plants with smaller leaves. Plants at high elevations and well away from the coast are even smaller (only 0.6 dm tall) with leaves only about 3 cm long. The high-elevation, ecotypic extremes are genetically fixed (T. A. Suttill and G. A. Allen 1992) but do not seem worthy of taxonomic recognition. Some individuals near Anchorage have maroon pollen sacs (compare LePage 23112 and LePage 23113, DAO).

Selected References

None.

Lower Taxa

None.

"reddish" is not a number. "maroon" is not a number. "purple" is not a number.

0.45 cm4.5 mm <br />0.0045 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (0.85 cm8.5 mm <br />0.0085 m <br />) +
acute;mostly acute;rounded +
James L. Reveal +
(A. Gray) Reveal +
whitish +  and yellowish +
tapering +  and decurrent +
Dodecatheon meadia var. macrocarpum +
crenulate +  and denticulate +
narrowly ovate;ovate +
puberulent +, glandular-pubescent +  and glabrous +
lanceolate +
0.2 cm2 mm <br />0.002 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
purple-flecked;green +
not keeled +  and tubular +
0.4 cm4 mm <br />0.004 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
1.4 cm14 mm <br />0.014 m <br /> (2 cm20 mm <br />0.02 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
, +, speckled with red or speckled with maroon +, reddish-brown +, tan +  and light-brown +
0.5 cm5 mm <br />0.005 m <br /> (1.4 cm14 mm <br />0.014 m <br />) +
glandular-pubescent +  and glabrous +
cylindric-ovoid +
0.3 cm3 mm <br />0.003 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
not obvious +
minute +
Alaskan shootingstar +
wrinkled +  and smooth +
yellow with red +  and maroon +
short-tubular +
B.C. +, Alaska +, Calif. +, Oreg. +  and Wash. +
0-1800 m +
0.17 cm1.7 mm <br />0.0017 m <br /> (0.27 cm2.7 mm <br />0.0027 m <br />) +
operculate +  and capsular +
Moist tidal flats, marshes, stream banks, slopes and cliffs mainly in meadows or grassy areas in or near conifer woodlands +
nectariferous +
2-15(-22)-flowered +
20 cm200 mm <br />0.2 m <br /> (35 cm350 mm <br />0.35 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
5 cm50 mm <br />0.05 m <br /> (20 cm200 mm <br />0.2 m <br />) +
1.5 cm15 mm <br />0.015 m <br /> (?) +  and 5 cm50 mm <br />0.05 m <br /> (?) +
pink to maroon speckles +
1.8 cm18 mm <br />0.018 m <br /> (2 cm20 mm <br />0.02 m <br />) +
magenta +  and lavender +
usually longer +
reflexed +  and spreading +
undulate +  and toothed +
membranous +
tenuinucellate +  and bitegmic +
5 cm50 mm <br />0.05 m <br /> (7 cm70 mm <br />0.07 m <br />) +
1 cm10 mm <br />0.01 m <br /> (5 cm50 mm <br />0.05 m <br />) +
white;pink or violet or magenta +
wingless +  and winged +
Flowering late spring–summer. +
purplish +, maroon +  and yellow +
puberulent +, glandular-pubescent +  and glabrous +
dark-brown +  and black +
alveolate +
globose +  and ovoid or quadrate +
toothlike +
Illustrated +
connate +  and distinct +
nearly +  and erect +
inflated +
not enlarged +
glandular-pubescent +  and glabrous +
Dodecatheon macrocarpum var. alaskanum +, Dodecatheon pauciflorum var. alaskanum +, Dodecatheon pulchellum subsp. alaskanum +, Dodecatheon pulchellum var. alaskanum +, Dodecatheon pulchellum subsp. macrocarpum +, Dodecatheon pulchellum subsp. superbum +  and Primula pauciflora var. macrocarpa +
Dodecatheon pulchellum var. macrocarpum +
Dodecatheon pulchellum +
variety +
maroon +  and dark purple or black +
0.07 cm0.7 mm <br />7.0e-4 m <br /> (0.36 cm3.6 mm <br />0.0036 m <br />) +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
1mm;3mm +
perennial +  and annual +
stoloniferous +  and rhizomatous +
45 cm450 mm <br />0.45 m <br /> (60 cm600 mm <br />0.6 m <br />) +