Eleocharis tenuis var. verrucosa

(Svenson) Svenson

Rhodora 41: 66. 1939.

IllustratedEndemic
Basionym: Eleocharis capitata var. verrucosa Svenson Rhodora 34: 202, plate 221, figs. 2, 14. 1932
Synonyms: Eleocharis verrucosa (Svenson) L. J. Harms
Treatment appears in FNA Volume 23. Treatment on page 88. Mentioned on page 84, 87.

Rhizomes (1–) 1.5–2 mm thick, hard, longer internodes 2 mm, scales decaying to fibers. Culms bluntly angled to smooth, seldom deeply sulcate, to 0.5 mm wide. Leaves: distal leaf-sheaths without tooth, or rarely with slender apical tooth to 0.2 mm. Flowers: perianth bristles absent. Achenes dark-brown, dark green, or yellow, coarsely (to finely) rugose at 10X, usually coarsely cancellate at 10–20X, 6–10 prominent depressions in each vertical series. Tubercles greatly depressed, rarely pyramidal. 2n = 20.


Phenology: Fruiting summer.
Habitat: Fresh, often calcareous, shores, wet woods, ditches
Elevation: 10–600 m

Distribution

V23 121-distribution-map.jpg

Ala., Ark., D.C., Ga., Ill., Ind., Iowa, Kans., Ky., La., Md., Mo., Nebr., Okla., Pa., S.Dak., Tenn., Tex., Va., W.Va., Wis.

Discussion

Occasionally plants of Eleocharis tenuis var. verrucosa are found mostly submerged.

Harms used a chromosome number as a basis for raising Eleocharis tenuis var. verrucosa to a species (L. J. Harms 1972), but he did not correlate chromosomes with morphology, and I have not located any of his voucher specimens. There are too few chromosome number reports for use in defining taxa in the E. tenuis complex (A. E. Schuyler 1977).

Selected References

None.

Lower Taxa

None.
yellow +, dark green +  and dark-brown +
terete;trigonous;biconvex +
color +  and texture +
green +, brown +, medium +, dark yellow +  and lemon yellow +
0.06 cm0.6 mm <br />6.0e-4 m <br /> (0.09 cm0.9 mm <br />9.0e-4 m <br />) +
prominent +  and evident +
alveolate +  and rugulose +
0.45mm;0.7mm +
0.08 cm0.8 mm <br />8.0e-4 m <br /> (0.18 cm1.8 mm <br />0.0018 m <br />) +
S. Galen Smith* +, Jeremy J. Bruhl* +, M. Socorro González-Elizondo* +  and Francis J. Menapace* +
(Svenson) Svenson +
flattened +
0 cm0 mm <br />0 m <br /> (6 cm60 mm <br />0.06 m <br />) +
shortened +
persistent +
enlarged +
Eleocharis capitata var. verrucosa +
glumaceous +  and foliaceous +
2-ranked +  and arranged +
ascending +  and appressed +
parallel +  and divergent +
terete +, rolled +  and plicate +
persistent +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.08 cm0.8 mm <br />8.0e-4 m <br />) +
5 cm50 mm <br />0.05 m <br /> (90 cm900 mm <br />0.9 m <br />) +
firm;soft +
0mm;0.5mm +
red +, stramineous +, green +  and dark red +
Ala. +, Ark. +, D.C. +, Ga. +, Ill. +, Ind. +, Iowa +, Kans. +, Ky. +, La. +, Md. +, Mo. +, Nebr. +, Okla. +, Pa. +, S.Dak. +, Tenn. +, Tex. +, Va. +, W.Va. +  and Wis. +
10–600 m +
membranous +  and thinly papery +
hypogynous +  and subtending +
biconvex +  and trigonous +
Fresh, often calcareous, shores, wet woods, ditches +
multi-ranked +, 2-ranked +, 3-ranked +  and alternate +
2 cm20 mm <br />0.02 m <br /> (?) +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
1 cm10 mm <br />0.01 m <br /> (?) +
flattened +
0 cm0 mm <br />0 m <br /> (6 cm60 mm <br />0.06 m <br />) +
with (1-)3-6(-30) bristles and/or scales +
Fruiting summer. +
2-3(-4)-carpellate +
3 (?) +  and 1 (?) +
stramineous;medium brown or red brown or blackish brown +
ascending +  and horizontal +
caudex-like +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
adventitious +
green +, brown +, medium +, dark yellow +  and lemon yellow +
decaying to fibers +  and persistent +
0.06 cm0.6 mm <br />6.0e-4 m <br /> (0.09 cm0.9 mm <br />9.0e-4 m <br />) +
basal +  and proximal +
prominent +  and evident +
rugulose +
trigonous;obpyriform +
0.5 cm5 mm <br />0.005 m <br /> (1 cm10 mm <br />0.01 m <br />) +
papery +  and finely coarsely fibrous +
0.45mm;0.7mm +
cylindric +
0 cm0 mm <br />0 m <br /> (0.02 cm0.2 mm <br />2.0e-4 m <br />) +
Illustrated +  and Endemic +
0.3 cm3 mm <br />0.003 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
1.5mm;2mm +
septate +, hollow +  and solid +
compressed +, terete +  and trigonous +
papillate +
Eleocharis verrucosa +
Eleocharis tenuis var. verrucosa +
Eleocharis tenuis +
variety +
0.02 cm0.2 mm <br />2.0e-4 m <br /> (0.09 cm0.9 mm <br />9.0e-4 m <br />) +
0 cm0 mm <br />0 m <br /> (0.02 cm0.2 mm <br />2.0e-4 m <br />) +
0.05mm;0.3mm +
pyramidal +  and depressed +
0.25mm +  and 0.4mm +
plant +  and mat-forming +