Potentilla brunnescens

Rydberg

Bull. Torrey Bot. Club 28: 173. 1901.

Endemic
Synonyms: Potentilla gracilis var. brunnescens (Rydberg) C. L. Hitchcock
Treatment appears in FNA Volume 9. Treatment on page 157. Mentioned on page 152, 158.

Glands conspicuous, usually golden. Stems ascending to nearly erect, (1.5–) 3–7 dm. Basal leaves sometimes 2-ranked, palmate, (5–) 10–30 cm; petiole (1.5–) 5–25 (–50) cm, long hairs sparse to abundant, appressed to spreading, 1–2 mm, ± stiff, short, crisped, and cottony hairs absent, glands ± abundant; leaflets (5–) 7–9, at tip of leaf axis, separate to slightly overlapping, largest ones oblanceolate-elliptic to narrowly obovate, (2–) 5–10 × 1–4 cm, margins flat to ± revolute, distal 3/4 to nearly whole length ± evenly incised (1/4–) 1/2–3/4+ to midvein, undivided medial blade 2–15 (–20) mm wide, teeth (5–) 7–10 per side, linear to broadly lanceolate, 3–15 mm, surfaces ± similar, abaxial paler, green, not glaucous, long hairs sparse to common, short-crisped hairs absent or sparse, cottony hairs absent, glands ± abundant. Cauline leaves 1–3 (–4). Inflorescences 12–30 (–70) -flowered. Pedicels 0.2–1 (–4) cm. Flowers: epicalyx bractlets lanceolate to elliptic, 3–6 × 1–2 mm, hairs sparse to common, loosely appressed to spreading, glands common to abundant; hypanthium 4–6 (–8) mm diam.; sepals 4–8 (–9) mm, apex acute to acuminate; petals 6–9 × 6–10 mm; filaments 1–2 mm, anthers 0.6–1 mm; carpels 30–60, styles filiform-tapered, papillate-swollen proximally, 1.5–2 mm. Achenes 1.5 mm.


Phenology: Flowering summer.
Habitat: Vernally moist rocky flats, sagebrush benchlands and slopes, usually basaltic
Elevation: 400–3100 m

Distribution

V9 230-distribution-map.jpg

Alaska, Colo., Idaho, Mont., Oreg., Utah, Wash., Wyo.

Discussion

The conspicuous golden glands, dark green color, and relatively broad leaflets make Potentilla brunnescens one of the better-defined species of sect. Graciles, though it has often been treated as a variety of P. gracilis. The glands, which are often on septate stalks, are particularly conspicuous on the epicalyx bractlets, hypanthium, and sepals, and at the junction of petiole and leaflets. The species occurs from the Blue Mountains of Oregon and Washington to western Montana, northern Utah, and north-central Colorado; collections from Alaska are probably not native.

As presented here, Potentilla brunnescens encompasses two morphological extremes that might merit varietal status. Leaflets of the more common and widespread extreme are incised about halfway to the midvein; in contrast, some populations in western Wyoming and adjacent Idaho have leaflets incised 3/4 or more to the midvein. The latter have often been assigned to P. gracilis var. elmeri while reserving P. gracilis var. brunnescens for the former; this is untenable because the type of P. brunnescens is the latter form.

Selected References

None.

Lower Taxa

None.

"dm" is not declared as a valid unit of measurement for this property.

... more about "Potentilla brunnescens"
stiff +  and weak +
green +  and paler +
blue-green +, green +  and grayish +
stiff +  and weak +
0.06 cm0.6 mm <br />6.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
Barbara Ertter +  and James L. Reveal +
Rydberg +
5 cm50 mm <br />0.05 m <br /> (10 cm100 mm <br />0.1 m <br />) +
10 cm100 mm <br />0.1 m <br /> (30 cm300 mm <br />0.3 m <br />) +
Graciles +
25 cm250 mm <br />0.25 m <br /> (30 cm300 mm <br />0.3 m <br />) +
less cordate or reniform +  and narrowly elliptic +
0.5 cm5 mm <br />0.005 m <br /> (25 cm250 mm <br />0.25 m <br />) +
coriaceous +
reduced +
free +  and distinct +
hairy +  and glabrous +
Alaska +, Colo. +, Idaho +, Mont. +, Oreg. +, Utah +, Wash. +  and Wyo. +
not +  and aggregated +
400–3100 m +
0.3 cm3 mm <br />0.003 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
lanceolate +  and elliptic +
0.1 cm1 mm <br />0.001 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
0.1 cm1 mm <br />0.001 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
0.8 cm8 mm <br />0.008 m <br /> (2 cm20 mm <br />0.02 m <br />) +
unisexual +  and bisexual +
not +  and aggregated +
deciduous +
on style scar +, hairy +  and glabrous +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.26 cm2.6 mm <br />0.0026 m <br />) +
conspicuous +
Vernally moist rocky flats, sagebrush benchlands and slopes, usually basaltic +
stoloniferous +  and short-rhizomatous +
blue-green +, green +  and grayish +
straight +
loosely appressed +  and spreading +
sparse +  and common +
short-crisped +  and crisped +
0.1 cm1 mm <br />0.001 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
0.7 cm7 mm <br />0.007 m <br /> (1 cm10 mm <br />0.01 m <br />) +
0.4 cm4 mm <br />0.004 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
12-30(-70)-flowered +
crassinucellate +
bipinnate +, odd-pinnate +, subpalmate +, palmate +  and ternate +
opposite +  and alternate +
stiff +  and weak +
distal +  and up-to-distal +
marcescent +
well-developed +
persistent +
cauline +  and basal +
reduced +
separate +  and slightly overlapping +
2 cm20 mm <br />0.02 m <br /> (5 cm50 mm <br />0.05 m <br />) +
5 cm50 mm <br />0.05 m <br /> (10 cm100 mm <br />0.1 m <br />) +
oblanceolate-elliptic +  and narrowly obovate +
1 cm10 mm <br />0.01 m <br /> (?) +  and 4 cm40 mm <br />0.04 m <br /> (?) +
linear +  and oblanceolate +
palmate +  and pinnate +
3/4 +  and length +
blue-green +, green +  and grayish +
stiff +  and weak +
1/4 +  and 1/2-3/4 +
flat +  and more or less revolute +
0.2 cm2 mm <br />0.002 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
blue-green +, green +  and grayish +
stiff +  and weak +
inferior +  and superior +
clustered +, biseriate +  and superposed +
1 cm10 mm <br />0.01 m <br /> (4 cm40 mm <br />0.04 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (1 cm10 mm <br />0.01 m <br />) +
1 cm10 mm <br />0.01 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
free +  and distinct +
0.6 cm6 mm <br />0.006 m <br /> (0.9 cm9 mm <br />0.009 m <br />) +
usually longer +
0.4 cm4 mm <br />0.004 m <br /> (1 cm10 mm <br />0.01 m <br />) +
0.6 cm6 mm <br />0.006 m <br /> (1 cm10 mm <br />0.01 m <br />) +
25 cm250 mm <br />0.25 m <br /> (50 cm500 mm <br />0.5 m <br />) +
5 cm50 mm <br />0.05 m <br /> (25 cm250 mm <br />0.25 m <br />) +
Flowering summer. +
adnate +, free +, connate +  and distinct +
subpinnate +, palmate +  and subpalmate +
not flagelliform +
2 cm20 mm <br />0.02 m <br /> (40 cm400 mm <br />0.4 m <br />) +
Bull. Torrey Bot. Club +
not arillate +
0.8 cm8 mm <br />0.008 m <br /> (0.9 cm9 mm <br />0.009 m <br />) +
persistent +
free +  and distinct +
erect +  and spreading +
lanceolate +  and broadly ovate or deltate +
0.4 cm4 mm <br />0.004 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
stiff +  and weak +
free +  and distinct +
not rooting +
reddish +  and green +
(1.5-)2-4 times basal leaves +
ascending +  and nearly erect +
1 +  and many +
not flagelliform +
palmate +  and pinnate +
linear;ovate +
deciduous +
distinct +
subapical +
papillate-swollen +  and filiform-tapered +
rough-thickened +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
blue-green +, green +  and grayish +
stiff +  and weak +
incised +, flat +  and more or less revolute +
Potentilla gracilis var. brunnescens +
not fleshy-thickened +
Potentilla brunnescens +
Potentilla sect. Graciles +
species +
blue-green +, green +  and grayish +
stiff +  and weak +
7 +  and 10 +
linear +  and broadly lanceolate +
0.3 cm3 mm <br />0.003 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
inconspicuous +
enlarged +
biennial +, annual +  and perennial +
herb +  and matted +