Potentilla drummondii

Lehmann

Nov. Stirp. Pug. 2: 9. 1830.

Common names: Drummond’s cinquefoil
Endemic
Synonyms: Potentilla cascadensis Rydberg P. drummondii var. cascadensis (Rydberg) Th. Wolf
Treatment appears in FNA Volume 9. Treatment on page 159. Mentioned on page 151, 160, 168, 170, 171, 172.

Glands usually absent or inconspicuous, rarely conspicuous, uncolored. Stems decumbent to nearly erect, 1.5–4.5 (–6) dm. Basal leaves sometimes 2-ranked, subpinnate to pinnate (proximal leaflets often doubled, distal leaflets confluent and/or decurrent), (4–) 10–25 cm; petiole 1–10 (–15) cm, long hairs absent or sparse to abundant, appressed, 1–1.5 mm, usually stiff, short, crisped, and cottony hairs usually absent, glands absent or (regionally) abundant; leaflets 5–9, on distal 1/10–1/2 (–3/4) of leaf axis, separate to ± overlapping, largest ones obovate to cuneate, (1–) 2–5 × (0.7–) 1–3 (–3.5) cm, margins flat, distal 1/2–3/4 unevenly, sometimes evenly, incised ± 1/2 to midvein (often with additional incisions nearly to midvein), undivided medial blade 3–15 mm wide, teeth 3–7 per side (sometimes secondarily toothed), linear to lanceolate, 3–10 mm, surfaces similar, green, not glaucous, long hairs nearly absent or sparse to common (sometimes restricted to abaxial veins), short, crisped, and cottony hairs usually absent, glands usually absent, sometimes regionally abundant. Cauline leaves 1–3. Inflorescences 3–15-flowered. Pedicels 1–3 (–6.5) cm. Flowers: epicalyx bractlets lanceolate to narrowly elliptic, 3–5 × 1–1.6 mm, hairs usually sparse, appressed to ascending, glands usually absent, sometimes regionally sparse to common; hypanthium (3–) 4–6 (–7) mm diam.; sepals 5–10 mm, apex acute to acuminate; petals (6–) 7–12 × 5–10 mm; filaments 2–3.5 mm, anthers 0.7–1 mm; carpels (20–) 25–40, styles filiform above papillate-swollen base, 2–3 mm. Achenes 1.5–2 mm. 2n = 64, 92, 96, 98–108.


Phenology: Flowering summer.
Habitat: Moist to dry meadows and adjacent slopes, in conifer woodlands, alpine tundra communities
Elevation: 300–3000 m

Distribution

V9 233-distribution-map.jpg

Alta., B.C., Alaska, Calif., Mont., Oreg., Wash.

Discussion

The present circumscription of Potentilla drummondii differs from that of B. Ertter (1992, 1993), which encompassed P. breweri and P. bruceae as varieties (see 28. P. bruceae discussion for details). The description above focuses on large, eglandular, subpinnate plants that occur in mountains from the central Sierra Nevada and northern Coast Ranges of California to the Olympic Peninsula of Washington, east to southern Alberta. At least some collections previously identified as P. glaucophylla (as P. diversifolia Lehmann) from the Kenai Peninsula in Alaska can be readily accommodated within this concept, as can collections from the Cabinet Mountains in Montana. The distinction between P. drummondii and subpalmate P. glaucophylla can be vague; in general, the proximalmost leaflets of P. glaucophylla are relatively small and often entire; in P. drummondii proximalmost leaflets are similar in size and dissection to other leaflets.

Pinnate-leaved regional variants merit further attention; in particular the smaller form described as Potentilla cascadensis, and glandular plants in the northern Coast Ranges of California. Relatively small plants forming uniform populations in the southern Sierra Nevada may represent stabilized hybrids with P. breweri.

The inclusion of Potentilla bruceae and P. drummondii in sect. Graciles differs from that of J. Clausen et al. (1940) and B. C. Johnston (1980, 1985), who allied both species with P. breweri in sect. Multijugae. Although subpalmate to subpinnate leaves are anomalous in primarily palmate sect. Graciles, doubled proximal leaflets and an indistinct transition between the rachis and axis of the pinnatisect terminal leaflet in these two species suggest that this state is derived from a palmate progenitor. Habit and petioles also are more compatible with sect. Graciles than sect. Multijugae.

Selected References

None.

Lower Taxa

None.

"dm" is not declared as a valid unit of measurement for this property.

... more about "Potentilla drummondii"
stiff +  and weak +
1.5mm +  and 2mm +
blue-green +, green +  and grayish +
stiff +  and weak +
0.07 cm0.7 mm <br />7.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
Barbara Ertter +  and James L. Reveal +
Lehmann +
subpinnate +  and pinnate +
4 cm40 mm <br />0.04 m <br /> (10 cm100 mm <br />0.1 m <br />) +
10 cm100 mm <br />0.1 m <br /> (25 cm250 mm <br />0.25 m <br />) +
papillate-swollen +
Graciles +
25 cm250 mm <br />0.25 m <br /> (30 cm300 mm <br />0.3 m <br />) +
less cordate or reniform +  and narrowly elliptic +
0.5 cm5 mm <br />0.005 m <br /> (25 cm250 mm <br />0.25 m <br />) +
coriaceous +
reduced +
free +  and distinct +
hairy +  and glabrous +
Drummond’s cinquefoil +
Alta. +, B.C. +, Alaska +, Calif. +, Mont. +, Oreg. +  and Wash. +
not +  and aggregated +
300–3000 m +
0.3 cm3 mm <br />0.003 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
lanceolate +  and narrowly elliptic +
0.1 cm1 mm <br />0.001 m <br /> (0.16 cm1.6 mm <br />0.0016 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
0.8 cm8 mm <br />0.008 m <br /> (2 cm20 mm <br />0.02 m <br />) +
unisexual +  and bisexual +
not +  and aggregated +
deciduous +
on style scar +, hairy +  and glabrous +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.26 cm2.6 mm <br />0.0026 m <br />) +
conspicuous +  and inconspicuous +
Moist to dry meadows and adjacent slopes, in conifer woodlands, alpine tundra communities +
stoloniferous +  and short-rhizomatous +
blue-green +, green +  and grayish +
straight +
appressed +  and ascending +
sparse +  and common +
crisped +
0.1 cm1 mm <br />0.001 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
0.7 cm7 mm <br />0.007 m <br /> (1 cm10 mm <br />0.01 m <br />) +
0.4 cm4 mm <br />0.004 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
crassinucellate +
bipinnate +, odd-pinnate +, subpalmate +, palmate +  and ternate +
opposite +  and alternate +
stiff +  and weak +
distal +  and up-to-distal +
marcescent +
well-developed +
persistent +
cauline +  and basal +
reduced +
separate +  and more or less overlapping +
1 cm10 mm <br />0.01 m <br /> (2 cm20 mm <br />0.02 m <br />) +
3 cm30 mm <br />0.03 m <br /> (3.5 cm35 mm <br />0.035 m <br />) +
2 cm20 mm <br />0.02 m <br /> (5 cm50 mm <br />0.05 m <br />) +
obovate +  and cuneate +
1 cm10 mm <br />0.01 m <br /> (?) +  and 3 cm30 mm <br />0.03 m <br /> (?) +
linear +  and oblanceolate +
palmate +  and pinnate +
blue-green +, green +  and grayish +
stiff +  and weak +
1/2 +  and 3/4 +
0.3 cm3 mm <br />0.003 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
blue-green +, green +  and grayish +
stiff +  and weak +
inferior +  and superior +
clustered +, biseriate +  and superposed +
3 cm30 mm <br />0.03 m <br /> (6.5 cm65 mm <br />0.065 m <br />) +
1 cm10 mm <br />0.01 m <br /> (3 cm30 mm <br />0.03 m <br />) +
0.6 cm6 mm <br />0.006 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
1 cm10 mm <br />0.01 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
free +  and distinct +
0.7 cm7 mm <br />0.007 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
usually longer +
0.4 cm4 mm <br />0.004 m <br /> (1 cm10 mm <br />0.01 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (1 cm10 mm <br />0.01 m <br />) +
10 cm100 mm <br />0.1 m <br /> (15 cm150 mm <br />0.15 m <br />) +
1 cm10 mm <br />0.01 m <br /> (10 cm100 mm <br />0.1 m <br />) +
Flowering summer. +
adnate +, free +, connate +  and distinct +
subpinnate +, palmate +  and subpalmate +
not flagelliform +
2 cm20 mm <br />0.02 m <br /> (40 cm400 mm <br />0.4 m <br />) +
Nov. Stirp. Pug. +
not arillate +
persistent +
free +  and distinct +
erect +  and spreading +
lanceolate +  and broadly ovate or deltate +
0.5 cm5 mm <br />0.005 m <br /> (1 cm10 mm <br />0.01 m <br />) +
stiff +  and weak +
free +  and distinct +
not rooting +
reddish +  and green +
(1.5-)2-4 times basal leaves +
decumbent +  and nearly erect +
1 +  and many +
not flagelliform +
palmate +  and pinnate +
linear;ovate +
deciduous +
distinct +
subapical +
filiform +
rough-thickened +
0.2 cm2 mm <br />0.002 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
stiff +  and weak +
not glaucous +
incised +, flat +  and more or less revolute +
Potentilla cascadensis +  and P. drummondii var. cascadensis +
not fleshy-thickened +
Potentilla drummondii +
Potentilla sect. Graciles +
species +
blue-green +, green +  and grayish +
stiff +  and weak +
3 +  and 7 +
linear +  and lanceolate +
0.3 cm3 mm <br />0.003 m <br /> (1 cm10 mm <br />0.01 m <br />) +
inconspicuous +
enlarged +
biennial +, annual +  and perennial +
herb +  and matted +