Potentilla sect. Subjugae

(Rydberg) A. Nelson in J. M. Coulter and A. Nelson

in J. M. Coulter and A. Nelson, New Man. Bot. Rocky Mt., 255. 1909.

Endemic
Basionym: Subjugae Rydberg Bull. Torrey Bot. Club 23: 397. 1896
Treatment appears in FNA Volume 9. Treatment on page 165. Mentioned on page 124, 126, 161, 166, 178, 181, 206.

Perennials, ± tufted, not stoloniferous; taproots not fleshy-thickened; vestiture of long, crisped, and/or cottony hairs, glands sparse to common, sometimes red. Stems ascending to nearly erect, not flagelliform, not rooting at nodes, lateral to persistent basal rosettes, (0.2–) 0.4–2.5 (–3.5) dm, lengths 2–3 times basal leaves. Leaves: basal not 2-ranked; cauline 0–3; primary leaves palmate with additional lateral pairs to pinnate (with distal leaflets distinct), (1–) 1.5–10 (–14) cm; petiole: long hairs spreading to tightly appressed, weak to stiff, glands absent or sparse; leaflets 5–7 (–9), on distal 1/10–1/3 (–1/2) of leaf axis, strongly overlapping or not, oblanceolate-oblong to obovate, margins scarcely to ± revolute, ± whole length evenly incised 1/2–3/4 to midvein, teeth 2–9 per side, surfaces ± to strongly dissimilar, abaxial white to grayish green, cottony hairs sparse to dense, adaxial green to grayish, not glaucous, long hairs ± stiff. Inflorescences (1–) 2–20 (–30) -flowered, usually cymose, open. Pedicels straight in fruit, 0.5–2 (–3) cm, proximal ± longer than distal. Flowers 5-merous; hypanthium 3–5 mm diam.; petals yellow, ± obcordate, 4–8 mm, equal to or longer than sepals, apex retuse; stamens ca. 20; styles subapical, filiform to filiform-tapered or filiform-columnar, papillate-swollen in proximal less than 1/5 if at all, 0.8–2 mm. Achenes smooth.

Distribution

w North America

Discussion

Species 2 (2 in the flora).

Section Subjugae is used primarily to accommodate the unique leaf dissection and vestiture of Potentilla subjuga, which commonly has five palmately arranged leaflets and one, sometimes two, additional pairs of lateral leaflets. Sometimes leaves are simply pinnate with only three terminal leaflets, and sufficient intergradation occurs with P. saximontana that it is also included in the section. This circumscription coincides with the initial concept of Rydberg and Nelson and contrasts with that of B. C. Johnston (1980, 1985).

Section Subjugae is restricted to subalpine and alpine habitats in the Rocky Mountains and adjacent ranges of western North America centered in Colorado. Variation within the section is complex and poorly understood, such that the two species treated here represent only the best expressed extremes with existing names. Other named components of uncertain placement include Potentilla lupina Rydberg, P. minutifolia Rydberg, P. rubricaulis var. nana Clements & E. G. Clements, P. rubripes Rydberg [= P. concinna var. rubripes (Rydberg) C. L. Hitchcock], and P. tenerrima Rydberg.

Selected References

None.

Key

1 Leaflets at tip of leaf axis (3–)5; styles 1.5–2 mm; stems (0.8–)1–2.5(–3.5) dm; inflorescences 3–20(–30)-flowered. Potentilla subjuga
1 Leaflets at tip of leaf axis 3; styles 0.8–1.5 mm; stems (0.2–)0.4–1(–1.5) dm; inflorescences (1–)2–5(–10)-flowered. Potentilla saximontana

"dm" is not declared as a valid unit of measurement for this property.

... more about "Potentilla sect. Subjugae"
white +  and grayish green +
green +  and grayish +
proximal +
Barbara Ertter +
(Rydberg) A. Nelson in J. M. Coulter and A. Nelson +
compound +  and simple +
Subjugae +
25 cm250 mm <br />0.25 m <br /> (30 cm300 mm <br />0.3 m <br />) +
less cordate or reniform +  and narrowly elliptic +
0.5 cm5 mm <br />0.005 m <br /> (25 cm250 mm <br />0.25 m <br />) +
reduced +
free +  and distinct +
hairy +  and glabrous +
w North America +
not +  and aggregated +
0.8 cm8 mm <br />0.008 m <br /> (2 cm20 mm <br />0.02 m <br />) +
unisexual +  and bisexual +
not +  and aggregated +
deciduous +
on style scar +, hairy +  and glabrous +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.26 cm2.6 mm <br />0.0026 m <br />) +
crisped +
stoloniferous +  and short-rhizomatous +
straight +
sparse +  and dense +
stiff +  and weak +
spreading +  and tightly appressed +
crisped +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
0.7 cm7 mm <br />0.007 m <br /> (1 cm10 mm <br />0.01 m <br />) +
0.3 cm3 mm <br />0.003 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
open +, cymose +  and (1-)2-20(-30)-flowered +
crassinucellate +
bipinnate +, odd-pinnate +, subpalmate +, palmate +  and ternate +
opposite +  and alternate +
marcescent +
well-developed +
persistent +
cauline +  and basal +
reduced +
not +  and overlapping +
oblanceolate-oblong +  and obovate +
linear +  and oblanceolate +
palmate +  and pinnate +
1/2 +  and 3/4 +
inferior +  and superior +
clustered +, biseriate +  and superposed +
2 cm20 mm <br />0.02 m <br /> (3 cm30 mm <br />0.03 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (2 cm20 mm <br />0.02 m <br />) +
free +  and distinct +
0.4 cm4 mm <br />0.004 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
adnate +, free +, connate +  and distinct +
10 cm100 mm <br />0.1 m <br /> (14 cm140 mm <br />0.14 m <br />) +
1.5 cm15 mm <br />0.015 m <br /> (10 cm100 mm <br />0.1 m <br />) +
in J. M. Coulter and A. Nelson, New Man. Bot. Rocky Mt., +
not arillate +
persistent +
free +  and distinct +
erect +  and spreading +
lanceolate +  and broadly ovate or deltate +
free +  and distinct +
not rooting +
reddish +  and green +
2-3 times basal leaves +
ascending +  and nearly erect +
1 +  and many +
not flagelliform +
palmate +  and pinnate +
deciduous +
distinct +
subapical +
papillate-swollen +, filiform +  and filiform-tapered or filiform-columnar +
rough-thickened +
0.08 cm0.8 mm <br />8.0e-4 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
not fleshy-thickened +
Potentilla sect. Subjugae +
Potentilla +
section +
2 +  and 9 +
inconspicuous +
enlarged +
biennial +, annual +  and perennial +
herb +  and matted +