Potentilla subjuga

Rydberg

Bull. Torrey Bot. Club 23: 397, plate 274. 1896.

Common names: Colorado cinquefoil
EndemicIllustrated
Synonyms: Potentilla osterhoutiana Th. Wolf
Treatment appears in FNA Volume 9. Treatment on page 166. Mentioned on page 161, 165, 178.

Stems (0.8–) 1–2.5 (–3.5) dm. Basal leaves usually palmate with additional lateral leaflets, sometimes pinnate, 3–10 (–14) cm; petiole 1.5–5 cm, vestiture seasonally dimorphic, long hairs abundant, spreading on first-formed leaves, tightly appressed to ascending on later-formed leaves, 1–2 mm, ± stiff (especially on later-formed leaves), cottony and crisped hairs usually absent, glands absent or sparse; leaflets (3–) 5 at tip of leaf axis plus 1 (–2) additional pair (s) separated from tip by 3–20 mm, on distal 1/10–1/3 (–1/2) of leaf axis, largest leaflets oblanceolate-oblong, (0.5–) 1.5–2.5 (–3) × 0.3–1 cm, ± whole margin incised 1/2–2/3 (–3/4) to midvein, teeth (2–) 4–9 per side, usually touching to strongly overlapping, sometimes separate, 2–6 mm, surfaces usually strongly dissimilar (less so on first-formed leaves), abaxial usually white, straight hairs ± abundant (mostly on veins), 1–2 mm, cottony or crisped/cottony hairs ± dense (sparser on first-formed leaves), glands absent or obscured, adaxial green (to grayish), straight hairs sparse to common, 0.5–1.5 mm, cottony and crisped hairs absent, glands sparse. Cauline leaves 1–3. Inflorescences 3–20 (–30) -flowered. Pedicels 0.5–2 cm (proximal to 3 cm). Flowers: epicalyx bractlets narrowly to broadly lanceolate, 2–5 (–6) × 1–1.5 mm; sepals 4–7 mm, apex acute to acuminate; petals 4–8 × 4–8 mm; filaments (0.5–) 1–2 mm, anthers 0.3–0.8 mm; carpels 15–30, styles filiform to filiform-tapered, ± papillate-swollen in less than proximal 1/5, 1.5–2 mm. Achenes 1.2–1.6 mm.


Phenology: Flowering summer.
Habitat: Alpine tundra and meadows, boulder piles, gravelly slopes, stabilized talus
Elevation: 3400–4000 m

Discussion

Potentilla subjuga is centered in the high mountains of Colorado and barely enters New Mexico in the Sangre de Cristo Mountains. Collections from Alberta also apparently belong to this species, but all known collections from Wyoming have been identified as different taxa, at least one currently undescribed. At its most distinctive, P. subjuga is easily recognized by its unique leaf division, with five palmate leaflets subtended by an additional pair (or two) of lateral leaflets. Southern populations, however, are more likely to have only three apical leaflets. The leaflets tend to be strongly bicolored with overlapping teeth, in contrast to most sympatric pinnate species. Petiole vestiture is also distinctive in being seasonally dimorphic, with long hairs on first-formed leaves spreading to ascending and those on later formed leaves tightly appressed, as well as more conspicuously verrucose. Unresolved infraspecific variation exists, and field observations suggest that P. subjuga readily hybridizes with sympatric species, creating a swarm of intermediate specimens.

Selected References

None.

Lower Taxa

None.

"dm" is not declared as a valid unit of measurement for this property.

... more about "Potentilla subjuga"
1.2mm +  and 1.6mm +
proximal +
0.03 cm0.3 mm <br />3.0e-4 m <br /> (0.08 cm0.8 mm <br />8.0e-4 m <br />) +
Barbara Ertter +
Rydberg +
10 cm100 mm <br />0.1 m <br /> (14 cm140 mm <br />0.14 m <br />) +
3 cm30 mm <br />0.03 m <br /> (10 cm100 mm <br />0.1 m <br />) +
Subjugae +
25 cm250 mm <br />0.25 m <br /> (30 cm300 mm <br />0.3 m <br />) +
less cordate or reniform +  and narrowly elliptic +
0.5 cm5 mm <br />0.005 m <br /> (25 cm250 mm <br />0.25 m <br />) +
reduced +
free +  and distinct +
hairy +  and glabrous +
Colorado cinquefoil +
Alta. +, Colo. +  and N.Mex. +
not +  and aggregated +
3400–4000 m +
0.5 cm5 mm <br />0.005 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
0.1 cm1 mm <br />0.001 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
0.1 cm1 mm <br />0.001 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
0.8 cm8 mm <br />0.008 m <br /> (2 cm20 mm <br />0.02 m <br />) +
unisexual +  and bisexual +
not +  and aggregated +
deciduous +
on style scar +, hairy +  and glabrous +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.26 cm2.6 mm <br />0.0026 m <br />) +
crisped +
Alpine tundra and meadows, boulder piles, gravelly slopes, stabilized talus +
stoloniferous +  and short-rhizomatous +
straight +
tightly appressed +  and ascending +
sparse +  and common +
crisped +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
0.7 cm7 mm <br />0.007 m <br /> (1 cm10 mm <br />0.01 m <br />) +
0.3 cm3 mm <br />0.003 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
3-20(-30)-flowered +
crassinucellate +
2.5 cm25 mm <br />0.025 m <br /> (3 cm30 mm <br />0.03 m <br />) +
1.5 cm15 mm <br />0.015 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
oblanceolate-oblong +
0.3 cm3 mm <br />0.003 m <br /> (1 cm10 mm <br />0.01 m <br />) +
bipinnate +, odd-pinnate +, subpalmate +, palmate +  and ternate +
opposite +  and alternate +
marcescent +
well-developed +
persistent +
cauline +  and basal +
reduced +
oblanceolate-oblong +  and obovate +
linear +  and oblanceolate +
palmate +  and pinnate +
1/2 +  and 2/3-3/4 +
inferior +  and superior +
clustered +, biseriate +  and superposed +
2 cm20 mm <br />0.02 m <br /> (3 cm30 mm <br />0.03 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (2 cm20 mm <br />0.02 m <br />) +
free +  and distinct +
0.4 cm4 mm <br />0.004 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
0.4 cm4 mm <br />0.004 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
0.4 cm4 mm <br />0.004 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
1.5 cm15 mm <br />0.015 m <br /> (5 cm50 mm <br />0.05 m <br />) +
Flowering summer. +
adnate +, free +, connate +  and distinct +
10 cm100 mm <br />0.1 m <br /> (14 cm140 mm <br />0.14 m <br />) +
1.5 cm15 mm <br />0.015 m <br /> (10 cm100 mm <br />0.1 m <br />) +
Bull. Torrey Bot. Club +
not arillate +
persistent +
free +  and distinct +
erect +  and spreading +
lanceolate +  and broadly ovate or deltate +
0.4 cm4 mm <br />0.004 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
Endemic +  and Illustrated +
free +  and distinct +
not rooting +
reddish +  and green +
2-3 times basal leaves +
ascending +  and nearly erect +
1 +  and many +
not flagelliform +
palmate +  and pinnate +
deciduous +
distinct +
subapical +
papillate-swollen +, filiform +  and filiform-tapered +
rough-thickened +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
separate +  and overlapping +
0.2 cm2 mm <br />0.002 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
Potentilla osterhoutiana +
not fleshy-thickened +
Potentilla subjuga +
Potentilla sect. Subjugae +
species +
4 +  and 9 +
inconspicuous +
enlarged +
biennial +, annual +  and perennial +
herb +  and matted +