Ranunculus pedatifidus var. affinis

(R. Brown) L. D. Benson

Amer. Midl. Naturalist 52: 355. 1954.

Endemic
Basionym: Ranunculus affinis R. Brown J. Voy. N.-W. Passage, Bot., 265. 1824
Synonyms: Ranunculus eastwoodianus L. D. Benson Ranunculus pedatifidus subsp. affinis (R. Brown) Hultén
Treatment appears in FNA Volume 3.

Stems erect, 6-33 (-46) cm, pilose or glabrous, each with 1-7 flowers. Roots slender, 0.4-1.2 mm thick. Basal leaves persistent, blades cordate or reniform in outline, pedately (5-) 7 (-9) -parted or divided, 0.8-3.8 × 1-4.8 cm, segments undivided or again lobed or parted, base truncate to cordate, margins never toothed, apices of segments acute. Flowers: pedicels pilose; receptacle canescent; sepals 4-6 × 3-5 mm, abaxially pilose, hairs colorless; petals (0-) 5-10, 7-10 × 5-9 mm; nectary scale glabrous. Heads of achenes cylindric, 7-15 × 5-8 mm; achenes 1.8-2.4 × 1.6-1.8 mm, finely canescent or glabrous; beak lanceolate or subulate, curved, 0.5-1 mm. 2n = 32, 48.


Phenology: Flowering late spring–summer (Jun–Aug).
Habitat: Dry rocky places on open arctic and alpine slopes and shores, moist grassland depressions, and open aspen woods
Elevation: 0-3700 m

Distribution

V3 1031-distribution-map.gif

Greenland, Alta., B.C., Man., Nfld. and Labr. (Nfld.), N.W.T., Ont., Que., Sask., Yukon, Alaska, Ariz., Colo., Mont., Utah, Wyo.

Discussion

Specimens of Ranunculus pedatifidus from southern Siberia, China, and central Asia, including the type specimen, are relatively small and slender, with all the main divisions of the leaf parted into narrow (1.5-2.5 mm wide) segments. American material, including the type of R. affinis, is more robust; the main divisions of the leaf are mostly undivided, and the ultimate segments are broader (2-4 mm). L. D. Benson (1954) referred all material from Asia and islands in the Bering Sea to var. pedatifidus. Other authors (P. N. Ovchinnikov 1970; E. Hultén 1968; A. I. Tolmatchew 1971), on the other hand, have limited typical R. pedatifidus to material from temperate Asia and have included material from northern Siberia in R. pedatifidus subsp. affinis. T. G. Tutin and J. R. Akeroyd (1993) included both R. pedatifidus and R. affinis in Flora Europaea, but, as noted below (under R. auricomus), Tutin and Akeroyd's descriptions do not match the American plants. Further comparisons of American and Eurasian material are needed.

Glabrous-fruited forms have been called Ranunculus pedatifidus var. leiocarpus (Trautvetter) Fernald; the variety was first described from Asia.

Selected References

None.

Lower Taxa

None.
not prolonged +
asymmetrically thick-lenticular +  and compressed-globose +
1.2-2 times as wide as thick +
0.18 cm1.8 mm <br />0.0018 m <br /> (0.24 cm2.4 mm <br />0.0024 m <br />) +
glabrous +  and canescent +
1.6mm;1.8mm +
Alan T. Whittemore +
(R. Brown) L. D. Benson +
truncate +  and cordate +
Ranunculus affinis +
subulate +  and lanceolate +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
0.8 cm8 mm <br />0.008 m <br /> (3.8 cm38 mm <br />0.038 m <br />) +
divided;(5-)7(-9)-parted;reniform;cordate +
1 cm10 mm <br />0.01 m <br /> (4.8 cm48 mm <br />0.048 m <br />) +
large +  and small +
globose;cylindric +
sometimes;sessile;nearly sessile +
1-50-flowered +
0 cm0 mm <br />0 m <br /> (25 cm250 mm <br />0.25 m <br />) +
Greenland +, Alta. +, B.C. +, Man. +, Nfld. and Labr. (Nfld.) +, N.W.T. +, Ont. +, Que. +, Sask. +, Yukon +, Alaska +, Ariz. +, Colo. +, Mont. +, Utah +  and Wyo. +
0-3700 m +
showy +  and inconspicuous +
sessile +  and aggregate +
cylindric;obovoid;cylindric;obovoid;globose;lenticular;discoid +
Dry rocky places on open arctic and alpine slopes and shores, moist grassland depressions, and open aspen woods +
0.7 cm7 mm <br />0.007 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
cylindric +
0.5 cm5 mm <br />0.005 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
axillary +  and terminal +
reniform +  and linear +
compound +  and simple +
whorled +, opposite +  and alternate +
cauline +  and basal +
parted +  and lobed +
palmate +  and pinnate +
inconspicuous +
distinct +
7mm;10mm +
conspicuous +
linear +  and orbiculate +
reduced +
0.1 cm1 mm <br />0.001 m <br /> (2.6 cm26 mm <br />0.026 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (0.9 cm9 mm <br />0.009 m <br />) +
Flowering late spring–summer (Jun–Aug). +
4 +  and 250 +
Amer. Midl. Naturalist +
slender +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.12 cm1.2 mm <br />0.0012 m <br />) +
not arillate +  and stalked +
again +  and undivided +
parted +  and lobed +
white;yellow;white;yellow;purple;sometimes;green +
deciduous +
distinct +
oblong +  and elliptic ovate or lanceolate +
0.1 cm1 mm <br />0.001 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
0.3 cm3 mm <br />0.003 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
(5-)10-many +
not bulbous-based +
33 cm330 mm <br />0.33 m <br /> (46 cm460 mm <br />0.46 m <br />) +
glabrous +  and pilose +
6 cm60 mm <br />0.06 m <br /> (33 cm330 mm <br />0.33 m <br />) +
persistent +
Ranunculus eastwoodianus +  and Ranunculus pedatifidus subsp. affinis +
Ranunculus pedatifidus var. affinis +
Ranunculus pedatifidus +
variety +
perennial +  and annual +
pubescent +, pilose +, sometimes +  and glabrous +
herbaceous +  and woody +