Rubus leucodermis

Douglas ex Torrey & A. Gray

Fl. N. Amer. 1: 454. 1840.

Synonyms: Rubus occidentalis subsp. leucodermis (Douglas ex Torrey & A. Gray) Focke
Treatment appears in FNA Volume 9. Treatment on page 44. Mentioned on page 31, 46.

Shrubs, 10–30 dm, armed. Stems biennial, ascending, often arching, glabrous or sparsely puberulent, eglandular or sparsely to densely short-stipitate-glandular, strongly pruinose; prickles moderate to dense, erect or curved, slender to stout, 4–8 mm, usually broad-based, sometimes narrow-based. Leaves deciduous, palmately compound or ternate; stipules filiform, 5–10 mm; petiole with hooked or erect prickles, glabrous, eglandular or sparsely to densely sessile to short-stipitate-glandular; lateral leaflets stalked, at least 2 larger ones, often with prickles, glabrous, eglandular or stipitate-glandular; leaflets 3–5 (–7), terminal ovate to lanceolate, 6–10 × 3–6 cm, base cordate to truncate, unlobed or shallowly 3-lobed, margins serrate to doubly serrate, apex acuminate, abaxial surface sometimes with prickles on midrib and some lateral-veins, glabrous to strongly white-tomentose, usually eglandular. Inflorescences axillary and terminal, (1–) 3–10 (–12) -flowered, flat-topped cymiform or umbelliform. Pedicels: prickles sparse to dense, hooked or erect, puberulent, eglandular or sparsely to densely sessile or short-stipitate-glandular. Flowers bisexual; petals erect, then ascending, white, oblong to oblanceolate, 3–6 (–8) mm; filaments laminar; ovaries pubescent, rarely glabrous. Fruits reddish purple to nearly black, depressed-globose to conic, 1–2 cm; drupelets 20–60, coherent, separating from torus. 2n = 14.

Distribution

V9 57-distribution-map.jpg

B.C., Alaska, Ariz., Calif., Idaho, Mont., N.Mex., Nev., Oreg., Utah, Wash., n Mexico

Discussion

Varieties 2 (2 in the flora).

Rubus leucodermis is similar to R. occidentalis, differing primarily by its leaflets having acuminate apices and lateral leaflets that are distinctly stalked. Prickles on the pedicels of R. occidentalis are erect; those of R. leucodermis tend to be hooked.

Selected References

None.

Key

1 Leaves abaxially white-tomentose; drupelets (20–)30–60. Rubus leucodermis var. leucodermis
1 Leaves abaxially glabrous, rarely sparsely hairy; drupelets 20–40. Rubus leucodermis var. nigerrimus

"thin" is not a number."dm" is not declared as a valid unit of measurement for this property.

... more about "Rubus leucodermis"
glabrous +  and strongly white-tomentose +
acuminate +
Lawrence A. Alice +, Douglas H. Goldman +, James A. Macklin +  and Gerry Moore +
Douglas ex Torrey & A. Gray +
compound +  and simple +
opposite +  and alternate +
cordate +  and truncate unlobed or shallowly 3-lobed +
reniform +  and orbiculate +
2 cm20 mm <br />0.02 m <br /> (30 cm300 mm <br />0.3 m <br />) +
herbaceous +  and more or less coriaceous +
not pruinose +
stipitate-glandular +, hairy +  and glabrous +
sparse to dense +
free +  and distinct +
hairy +  and glabrous +
B.C. +, Alaska +, Ariz. +, Calif. +, Idaho +, Mont. +, N.Mex. +, Nev. +, Oreg. +, Utah +, Wash. +  and n Mexico +
not +  and aggregated +
0.5 cm5 mm <br />0.005 m <br /> (8 cm80 mm <br />0.08 m <br />) +
not +  and aggregated +
without torus +  and separating +
reddish purple +  and nearly black +
coherent +
hairy +  and glabrous +
depressed-globose +  and conic +
1 cm10 mm <br />0.01 m <br /> (2 cm20 mm <br />0.02 m <br />) +
dryish +  and fleshy +
eglandular or +  and sparsely densely glandular +
0.3 cm3 mm <br />0.003 m <br /> (1 cm10 mm <br />0.01 m <br />) +
glabrous or +  and sparsely densely pubescent +
flat +  and hemispheric +
umbelliform +, cymiform +, flat-topped +  and (1-)3-10(-12)-flowered +
glabrous or +  and sparsely densely pubescent +
crassinucellate +
eglandular +  and stalked +
stipitate-glandular +  and glabrous +
ternate +  and compound +
deciduous +
6 cm60 mm <br />0.06 m <br /> (10 cm100 mm <br />0.1 m <br />) +
ovate +  and lanceolate +
1.7 cm17 mm <br />0.017 m <br /> (15 cm150 mm <br />0.15 m <br />) +
3 cm30 mm <br />0.03 m <br /> (?) +  and 6 cm60 mm <br />0.06 m <br /> (?) +
revolute +  and finely coarsely crenate +
inferior +  and superior +
glabrous +  and pubescent +
clustered +, biseriate +  and superposed +
collateral +  and apical +
1 +  and 2 +
eglandular +, armed +  and unarmed +
stipitate-glandular +, glabrous or +  and sparsely densely hairy +
0 (?) +  and 5 (?) +
0.6 cm6 mm <br />0.006 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
free +  and distinct +
ascending +  and erect +
oblong;oblanceolate +
0.3 cm3 mm <br />0.003 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
eglandular or +  and sparsely densely sessile +
short-stipitate-glandular +  and glabrous +
adnate +, free +, connate +  and distinct +
eglandular or +  and sparsely densely sessile +
sparse +  and dense +
short-stipitate-glandular +  and puberulent +
sparse to dense +
hooked +  and broad-based +
slender +  and stout +
0.4 cm4 mm <br />0.004 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
Fl. N. Amer. +
not arillate +
eglandular +, armed +  and unarmed +
persistent +
free +  and distinct +
reflexed +  and spreading +
stipitate-glandular +, hairy +  and glabrous +
lanceolate +  and long-caudate +
free +  and distinct +
shorter to longer +
pruinose +
scrambling +  and mounding +
decumbent +  and creeping +
arching +  and ascending +
short-stipitate-glandular;puberulent;glabrous +
1 +  and several +
angled +  and terete +
palmate +  and pinnate +
adnate +  and free +
0.5 cm5 mm <br />0.005 m <br /> (1 cm10 mm <br />0.01 m <br />) +
persistent +
distinct +
clavate +
slender +
Rubus occidentalis subsp. leucodermis +
Rubus leucodermis +
species +
inconspicuous +
convex +  and conic +
enlarged +  and small +
fibrous +  and woody +