Rubus leucodermis var. nigerrimus

(Greene) H. St. John

Fl. S.E. Washington, 213. 1937.

Common names: Dark or northwest raspberry
Conservation concernEndemic
Basionym: Melanobatus nigerrimus Greene Leafl. Bot. Observ. Crit. 1: 244. 1906,
Synonyms: R. nigerrimus (Greene) Rydberg
Treatment appears in FNA Volume 9. Treatment on page 45. Mentioned on page 44.

Shrubs, 10–20 dm, densely armed. Leaves abaxially glabrous, rarely sparsely hairy. Inflorescences 1–5-flowered. Flowers: petals oblong, 4–7 mm. Fruits: drupelets 20–40, glabrous, rarely puberulent.


Phenology: Flowering Apr–Aug.
Habitat: Moist to dry shrubby thickets and open forests in canyons
Elevation: 200–700 m

Discussion

Of conservation concern.

Variety nigerrimus can be distinguished from the typical variety by its essentially glabrous foliage. This variety is endemic to southeastern Washington near the Snake River in Garfield and Whitman counties. M. E. Peck (1941) reported it (as R. hesperius) from Oregon; no collections from there have been seen as part of this review. The taxonomic status of this variant remains questionable as characters such as pubescence are not discrete and may reflect the effect of environment.

Selected References

None.

Lower Taxa

None.

"thin" is not a number."dm" is not declared as a valid unit of measurement for this property.

glabrous +  and strongly white-tomentose +
acuminate +
Lawrence A. Alice +, Douglas H. Goldman +, James A. Macklin +  and Gerry Moore +
(Greene) H. St. John +
compound +  and simple +
opposite +  and alternate +
cordate +  and truncate unlobed or shallowly 3-lobed +
Melanobatus nigerrimus +
reniform +  and orbiculate +
2 cm20 mm <br />0.02 m <br /> (30 cm300 mm <br />0.3 m <br />) +
herbaceous +  and more or less coriaceous +
not pruinose +
stipitate-glandular +, hairy +  and glabrous +
sparse to dense +
free +  and distinct +
hairy +  and glabrous +
Dark or northwest raspberry +
not +  and aggregated +
puberulent +  and glabrous +
200–700 m +
0.5 cm5 mm <br />0.005 m <br /> (8 cm80 mm <br />0.08 m <br />) +
not +  and aggregated +
without torus +  and separating +
reddish purple +  and nearly black +
coherent +
hairy +  and glabrous +
depressed-globose +  and conic +
1 cm10 mm <br />0.01 m <br /> (2 cm20 mm <br />0.02 m <br />) +
dryish +  and fleshy +
Moist to dry shrubby thickets and open forests in canyons +
eglandular or +  and sparsely densely glandular +
0.3 cm3 mm <br />0.003 m <br /> (1 cm10 mm <br />0.01 m <br />) +
glabrous or +  and sparsely densely pubescent +
flat +  and hemispheric +
glabrous or +  and sparsely densely pubescent +
crassinucellate +
eglandular +  and stalked +
stipitate-glandular +  and glabrous +
ternate +  and compound +
deciduous +
hairy +  and glabrous +
6 cm60 mm <br />0.06 m <br /> (10 cm100 mm <br />0.1 m <br />) +
ovate +  and lanceolate +
1.7 cm17 mm <br />0.017 m <br /> (15 cm150 mm <br />0.15 m <br />) +
3 cm30 mm <br />0.03 m <br /> (?) +  and 6 cm60 mm <br />0.06 m <br /> (?) +
revolute +  and finely coarsely crenate +
inferior +  and superior +
glabrous +  and pubescent +
clustered +, biseriate +  and superposed +
collateral +  and apical +
1 +  and 2 +
eglandular +, armed +  and unarmed +
stipitate-glandular +, glabrous or +  and sparsely densely hairy +
0 (?) +  and 5 (?) +
0.6 cm6 mm <br />0.006 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
free +  and distinct +
ascending +  and erect +
0.4 cm4 mm <br />0.004 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
eglandular or +  and sparsely densely sessile +
short-stipitate-glandular +  and glabrous +
Flowering Apr–Aug. +
adnate +, free +, connate +  and distinct +
eglandular or +  and sparsely densely sessile +
sparse +  and dense +
short-stipitate-glandular +  and puberulent +
sparse to dense +
hooked +  and broad-based +
slender +  and stout +
0.4 cm4 mm <br />0.004 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
Fl. S.E. Washington, +
not arillate +
eglandular +, armed +  and unarmed +
persistent +
free +  and distinct +
reflexed +  and spreading +
stipitate-glandular +, hairy +  and glabrous +
lanceolate +  and long-caudate +
Conservation concern +  and Endemic +
free +  and distinct +
shorter to longer +
pruinose +
scrambling +  and mounding +
decumbent +  and creeping +
arching +  and ascending +
short-stipitate-glandular;puberulent;glabrous +
1 +  and several +
angled +  and terete +
palmate +  and pinnate +
adnate +  and free +
0.5 cm5 mm <br />0.005 m <br /> (1 cm10 mm <br />0.01 m <br />) +
persistent +
distinct +
clavate +
slender +
R. nigerrimus +
Rubus leucodermis var. nigerrimus +
Rubus leucodermis +
variety +
inconspicuous +
convex +  and conic +
enlarged +  and small +
fibrous +  and woody +