Mentioned on page 6. Plants annual, monoecious, climbing; stems glabrous or sparsely puberulent; roots fibrous; tendrils 2-branched. Leaves: blade broadly to
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× 2–6 mm, margins deeply 1- or multilobed, usually glandular, teeth lobelike, (3 or) 4 or 5 (or 6) per side on distal 1/2 of blades, some multi-serrate
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with sparse or moderately dense stipitate-glandular hairs on ribs, less densely pubescent or glabrous in sulci [rarely entirely glabrous]; ribs 5, round
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also sometimes glandular puberulent on distal parts. Stems slender, unbranched or branched from base, 10–60 cm. Leaves in basal rosette and/or cauline, often
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acuminate, glabrous to slightly glandular; peduncle and rachis glandular-pubescent. Flowers yellowish green, small; pedicel filiform, 4–12 mm, glandular-puberulent;
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12, 14, 57, 80, 82. Herbs, perennial; caude× woody or herbaceous. Stems erect, glabrous or puberulent. Leaves basal and cauline, cauline smaller, opposite
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ciliate, apex rounded to acute, surfaces: proximals glabrous, distals glandular-puberulent or glandular-pubescent. Pedicels 1–4 (–7 on proximal) mm in fruit
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stipitate-glandular; bracts 2, ovatelanceolate, 16–21 × 4–6 mm, margins glandular, surfaces tomentose to puberulent or glabrous, glandular or eglandular
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black; tips not or weakly spinescent; glabrous or sparsely to ± densely puberulent. Leaves: stipules absent; petiole base ± persistent, not or only slightly
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entire or glandular-serrate, eglandular or gland-tipped, surfaces glabrous, eglandular; petiole and rachis with pricklets, glabrous, rarely puberulent, eglandular;
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flared, 4–6 mm, margins entire or unevenly serrate, sessile-glandular, surfaces rugose, glabrous or pubescent, glandular; petiole and rachis with pricklets
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irregularly glandular-serrate, surfaces glabrous or pubescent, eglandular. Flowers 3–4.5 cm diam.; hypanthium globose, 5–7 × 5–7 mm, stipitate-glandular or setose
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pale pinkish or pale-yellow; stigmas (4–) 6 or 7 (–10). Schizocarps 4–7 mm diam.; mericarps usually (4–) 6 or 7 (–10), (1.5–) 2–2.5 mm, glabrous, sides usually
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1–2 m. Stems erect to spreading or prostrate, finely puberulent, sometimes glabrous except for prickles, not glandular; spines at nodes 1–3, 3–12 mm; prickles
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architecture or coloration or relief","fruit development","fruit pubescence","fruit shape","inflorescence architecture or arrangement or growth form","inflorescence
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rounded and shallowly 2-fid, abaxial limb glabrous or sparsely bearded. Styles glabrous. Anthers included, glabrous. Capsules included, 4–5 mm. Phenology:
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architecture","face architecture or function or pubescence","head architecture or arrangement","head architecture or shape","head arrangement","involucre
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twigs yellow-green, reddish green, or reddish-brown, terete to somewhat angled, finely puberulent or, rarely, glabrous. Leaf-blades green, usually oblanceolate
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inconspicuously glandular, apex usually acute, sometimes obtuse, surfaces usually glabrous, sometimes puberulent. Inflorescences solitary flowers or 2-flowered
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surfaces glabrous or sparsely puberulent; petals 25+, white or pink, elliptic to narrowly elliptic, 8–11 mm; ovaries glabrous. Drupes reddish, subglobose, 10–15
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