Carex sect. Granulares

(O. Lang) Mackenzie in N. L. Britton et al.

in N. L. Britton et al., N. Amer. Fl. 18: 260. 1935.

Basionym: Granulares O. Lang Linnaea 24: 582. 1851
Treatment appears in FNA Volume 23.
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Plants cespitose or not, short to long rhizomatous, sometimes inconspicuously rhizomatous. Culms sometimes solitary, brown at base. Leaves: basal sheaths not fibrous; sheath fronts membranous; blades M-shaped in cross-section when young, adaxial side of blade with 2 lateral-veins more prominent than midvein, widest leaves not more than 10 mm wide, glabrous. Inflorescences racemose, with (2–) 3–6 spikes; proximal nonbasal bracts leaflike, long-sheathing, sheath more than 4 mm, longer than diameter of stem; lateral spikes pistillate or androgynous, rarely distal 1–3 spikes staminate, sometimes some basal, pedunculate, prophyllate; terminal spike staminate. Proximal pistillate scales with apex acute, acuminate, or short-awned. Perigynia ascending to spreading, minute redbrown or yellowish streaks and dots, distinctly veined, sessile, ellipsoid or rhomboid to broadly ovoid, obovoid, or subglobose, obscurely trigonous to round in cross-section, base rounded, apex rounded, abruptly beaked, glabrous; beak 0.1–0.9 mm, orifice entire to minutely bidentulate, teeth 0–0.3 mm. Stigmas 3. Achenes trigonous, smaller than bodies of perigynia; style deciduous.

Distribution

Temperate and subtropical regions of North America, s Mexico, Central America (Guatemala)

Discussion

Species 6 (4 in the flora).

Carex sect. Granulares is a monophyletic section diagnosed by at least three apormorphies: red-brown or yellowish cells scattered in the epidermis of the perigynia, often 25 or more perigynia per well-developed lateral spike, and perigynia loosely enveloping the achenes (R. F. C. Naczi 1992, 1997). Recent phylogenetic analyses indicate section Granulares is the sister group of a clade composed of sections Careyanae and Griseae (R. F. C. Naczi 1992).

Selected References

None.

Key

1 Plants with short or inconspicuous rhizomes; culms in tufts; terminal spike and distal lateral spike usually overlapping; proximal spikes usually arising from distal 1/2 of culms. > 2
1 Plants with long-creeping rhizomes; culms mostly solitary; terminal spike and distal lateral spike (unless staminate) usually separated; proximal spikes usually arising from proximal 1/2 of culms. > 3
2 Leaves green, not glaucous; longest bract blade (per plant) of distal lateral spike 1.6–4.6(–7.1) cm; ligule of proximal bract 0.5–6.5 mm; perigynia (1.6–)1.9–3 times as long as thick. Carex gholsonii
2 Leaves not green, usually glaucous; longest bract blade (per plant) of distal lateral spike 4.1–15.8 cm; ligule of proximal bract (2–)3–17.5(–26) mm; perigynia 1.4–2.2(–2.4) times as long as thick. Carex granularis
3 Staminate scales with apex acute to awned; widest leaves 2.8–8.3 mm wide; perigynium beak 0.3–0.9 mm. Carex microdonta
3 Staminate scales with apex rounded to obtuse; widest leaves 1.8–3(–4.4) mm wide; perigynium beak 0.1–0.3 mm. Carex crawei

"shortened" is not a number.

... more about "Carex sect. Granulares"
trigonous +
smaller +
rounded;acute +
Theodore S. Cochrane +  and Robert F. C. Naczi +
(O. Lang) Mackenzie in N. L. Britton et al. +
rounded +
Granulares +
0.01 cm0.1 mm <br />1.0e-4 m <br /> (0.09 cm0.9 mm <br />9.0e-4 m <br />) +
m--shaped +
0 cm0 mm <br />0 m <br /> (2 cm20 mm <br />0.02 m <br />) +
glumaceous +  and foliaceous +
2-ranked +  and arranged +
ascending +  and appressed +
scale-like +  and leaflike +
parallel +  and divergent +
terete +, rolled +  and plicate +
round +  and trigonous +
Temperate and subtropical regions of North America +, s Mexico +  and Central America (Guatemala) +
open +, pistillate +  and staminate +
hypogynous +  and subtending +
biconvex +  and trigonous +
multi-ranked +, 2-ranked +, 3-ranked +  and alternate +
basal +  and cauline +
entire;minutely bidentulate +
with (1-)3-6(-30) bristles and/or scales +
yellowish +, redbrown +  and minute +
ascending +  and spreading +
2-3(-4)-carpellate +
3 (?) +  and 1 (?) +
long-sheathing +  and leaflike +
short-awned +  and acuminate +
in N. L. Britton et al., N. Amer. Fl. +
adventitious +
basal +  and proximal +
2-keeled +
cylindric +
prophyllate +, pedunculate +  and staminate +
1 +  and 3 +
septate +, hollow +  and solid +
compressed +, terete +  and trigonous +
papillate +
sessile +  and veined +
rhomboid;broadly ovoid obovoid or subglobose obscurely trigonous +
deciduous +
2-3(-4)-fid +
Carex sect. Granulares +
section +
0 cm0 mm <br />0 m <br /> (0.03 cm0.3 mm <br />3.0e-4 m <br />) +
plant +, not +  and cespitose +