Carex aquatilis var. substricta

Kükenthal in H. G. A. Engler

in H. G. A. Engler, Pflanzenr. 38[IV,20]: 309. 1909.

Common names: Carex subétroit
IllustratedEndemic
Synonyms: Carex substricta (Kükenthal) Mackenzie
Treatment appears in FNA Volume 23. Treatment on page 398. Mentioned on page 397.

Culms 35–100 cm, angles glabrous or scabrous. Leaves: basal sheaths leafless, usually redbrown; blades amphistomic, 5–8 mm wide, papillose on both surfaces. Inflorescences: peduncle of proximal spike not more than 4 cm; proximal bract longer than inflorescence, 4–8 mm wide. Spikes erect; proximal 2–6 spikes pistillate, 4–6 cm × 4–7 mm, base cuneate or attenuate; terminal 1–3 spikes staminate. Pistillate scales pale redbrown with narrow redbrown marginand broad pale midvein. Perigynia pale-brown, veinless, obovoid, 2.6–3.2 × 1.5–2.2 mm, apex rounded; beak pale-brown, thickened, not more than 0.2 mm. 2n = 76, 77.


Phenology: Fruiting Jul–Aug.
Habitat: Marshes, shallow water along shores, usually in neutral or calcareous substrates
Elevation: 0–1000 m

Distribution

V23 712-distribution-map.jpg

Man., N.B., N.S., Ont., Que., Conn., Ill., Ind., Iowa, Kans., Maine, Mass., Mich., Minn., Mo., Nebr., N.J., N.Y., N.Dak., Ohio, Pa., S.Dak., Vt., Va., Wis.

Discussion

Populations of Carex aquatilis var. substricta from Minnesota to Massachusetts tend to have very pale scales with a very broad, pale brown midvein, distinctly obovoid perigynia, and broad pistillate spikes that are rarely staminate at the apex. These populations tend to occur on neutral or calcareous substrates (fens) rather than the more acidic marshes favored by var. aquatilis. The two varieties intergrade where they come into contact.

Selected References

None.

Lower Taxa

None.

"not undefined" is not a number.

biconvex +
not constricted +
scabrous +  and glabrous +
papillose +
Lisa A. Standley +, Jacques Cayouette +  and Leo Bruederle +
Kükenthal in H. G. A. Engler +
shortened +
not +  and fibrous +
attenuate +  and cuneate +
bidentate +  and emarginate +
papillose +
m--shaped +  and v--shaped +
5mm;8mm +
glumaceous +  and foliaceous +
2-ranked +  and arranged +
ascending +  and appressed +
scale-like +  and leaflike +
parallel +  and divergent +
terete +, rolled +  and plicate +
Carex subétroit +
redbrown +  and brown +
round +  and trigonous +
35 cm350 mm <br />0.35 m <br /> (100 cm1,000 mm <br />1 m <br />) +
prophyllate +, subsessile +, pedunculate +  and staminate +
Man. +, N.B. +, N.S. +, Ont. +, Que. +, Conn. +, Ill. +, Ind. +, Iowa +, Kans. +, Maine +, Mass. +, Mich. +, Minn. +, Mo. +, Nebr. +, N.J. +, N.Y. +, N.Dak. +, Ohio +, Pa. +, S.Dak. +, Vt. +, Va. +  and Wis. +
0–1000 m +
open +, pistillate +  and staminate +
hypogynous +  and subtending +
biconvex +  and trigonous +
Marshes, shallow water along shores, usually in neutral or calcareous substrates +
multi-ranked +, 2-ranked +, 3-ranked +  and alternate +
basal +  and cauline +
pale +  and redbrown +
with (1-)3-6(-30) bristles and/or scales +
0.26 cm2.6 mm <br />0.0026 m <br /> (0.32 cm3.2 mm <br />0.0032 m <br />) +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.22 cm2.2 mm <br />0.0022 m <br />) +
Fruiting Jul–Aug. +
2-3(-4)-carpellate +
3 (?) +  and 1 (?) +
short-sheathing +, sheathless +  and leaflike +
0.4 cm4 mm <br />0.004 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
in H. G. A. Engler, Pflanzenr. +
adventitious +
pale redbrown +
basal +  and proximal +
2-keeled +
ladder-fibrillose +  and veined +
cylindric +
Illustrated +  and Endemic +
staminate +  and pistillate +
4 cm40 mm <br />0.04 m <br /> (6 cm60 mm <br />0.06 m <br />) +
0.4 cm4 mm <br />0.004 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
indistinct +
1 (?) +  and 3 (?) +
septate +, hollow +  and solid +
compressed +, terete +  and trigonous +
papillate +
deciduous +
2-3(-4)-fid +
Carex substricta +
Carex aquatilis var. substricta +
Carex aquatilis +
variety +
, +  and staminate +
plant +  and not cespitose +