Persicaria sect. Echinocaulon

(Meisner) H. Gross

Bull. Acad. Int. Géogr. Bot. 4: 27. 1913.

Basionym: Polygonum sect. Echinocaulon Meisner in N. Wallich, Pl. Asiat. Rar. 3: 58. 1832
Treatment appears in FNA Volume 5. Treatment on page 575. Mentioned on page 574.

Stems usually scandent or, rarely, ascending or erect, simple or branched, with recurved prickles. Leaves: ocrea chartaceous or foliaceous, margins eciliate or ciliate; petiole, if present, rarely winged, not auriculate; blade without dark triangular or lunate blotch adaxially, broadly lanceolate or elliptic to triangular, base truncate to sagittate to cordate, margins entire or hastately lobed. Inflorescences terminal and axillary, spikelike, paniclelike, or capitate, uninterrupted or, rarely, interrupted proximally. Flowers homostylous, articulation swollen or not; perianth campanulate; tepals 4–5, connate 1/4–1/2 their length; stamens 5–8; styles deciduous, 2–3, included or, rarely, exserted, erect or spreading.

Distribution

e North America, South America, Asia, se Africa, e Australia

Discussion

Species 21 (5 in the flora).

Key

1 Ocreae, at least some, foliaceous, green; perianths fleshy and blue in fruit Persicaria perfoliata
1 Ocreae chartaceous, brownish; perianths not fleshy and blue in fruit > 2
2 Ocreae margins truncate, ciliate with bristles 2-4 mm Persicaria bungeana
2 Ocreae margins oblique, eciliate or ciliate with bristles to 2.5 mm > 3
3 Leaf blades broadly hastate to hastate-cordate or triangular; tepals 4 Persicaria arifolia
3 Leaf blades linear-lanceolate to oblong; tepals 5 > 4
4 Peduncles usually glabrous, sometimes with retrorse prickles proximally; leaves petiolate; bases of leaf blades sagittate to cordate; stamens 8 Persicaria sagittata
4 Peduncles usually stipitate-glandular; leaves sessile or subsessile; bases of leaf blades cordate to truncate or cuneate; stamens 5 Persicaria meisneriana

"/2" is not declared as a valid unit of measurement for this property.

dark-brown;black +
exserted +  and included +
spheroidal +, 2-3-gonous +, biconvex +  and discoid +
red +, pink +  and yellow +
elliptic +  and ovate +
not +  and swollen +
Harold R. Hinds† +  and Craig C. Freeman +
(Meisner) H. Gross +
truncate +  and sagittate +
Polygonum sect. Echinocaulon +
elliptic;triangular +
lunate +, triangular +  and dark +
foliaceous +  and perfoliate +
scale-like +
just below the soil surface or and above the soil surface +  and spreading +
e North America +, South America +, Asia +, se Africa +  and e Australia +
adnate +, connate +  and distinct +
disarticulating +  and solid +
decumbent to erect +  and prostrate +
unwinged +  and winged +
black +, red +, brown +  and yellowish +
exserted +  and included +
pubescent +  and glabrous +
spheroidal +, 4-gonous +, lenticular +, biconvex +, discoid +, 3-gonous +  and 2-gonous +
interrupted +, , +  and uninterrupted +
paniclelike +  and spikelike +
nonmembranous +
deciduous +
not auriculate +
ciliate +  and eciliate +
lobed +  and entire +
swollen +
reddish +, brown +  and tan +
disintegrating +
deciduous +  and persistent +
scabrous +  and variously pubescent +
coriaceous +
anatropous +, , +  and orthotropous +
accrescent +
spreading;erect +
purple;red;purple;red;roseate;greenish white;white +
campanulate +
nonaccrescent +  and accrescent +
(2-)3(-4)-carpellate +
Bull. Acad. Int. Géogr. Bot. +
park1986a +, park1987a +  and park1988a +
slender;stout +
5 +  and 8 +
branched +  and simple +
, +  and scandent +
erect +  and ascending +
pubescent +  and glabrous +
penicillate +, fimbriate +  and capitate +
deciduous +  and persistent +
2-lobed +  and cylindric +
coriaceous +  and membranous +
deciduous +
connate +  and distinct +
spreading;erect +
exserted +, , +  and included +
spreading;erect +
Persicaria sect. Echinocaulon +
Persicaria +
section +
1/4 +  and 1/2 +
4 +  and 5 +
lobed +  and laciniate +
coriaceous +
persistent +
membranous +
pubescent +  and glabrous +
stoloniferous +, rhizomatous +, fibrous-rooted +  and taprooted +
annual +  and perennial +
12 +, 11 +  and 10 +