Potentilla gracilis var. elmeri

(Rydberg) Jepson

Fl. Calif. 2: 189. 1936.

Common names: Elmer’s or combleaf cinquefoil
EndemicIllustrated
Basionyms: Potentilla elmeri Rydberg in N. L. Britton et al., N. Amer. Fl. 22: 315. 1908
Synonyms: P. bakeri Rydberg P. flabelliformis var. inyoensis Jepson P. pectinisecta Rydberg P. pectinisecta var. comosa (Rydberg) Soják
Treatment appears in FNA Volume 9. Treatment on page 156. Mentioned on page 154, 158.

Stems (1.5–) 2–6 (–8) dm. Basal leaves: petiole 2–15 (–25) cm, long hairs common to dense, usually tightly appressed, sometimes ascending to spreading, 0.5–2.5 mm, stiff, crisped hairs absent or sparse, cottony hairs absent, glands absent or sparse; leaflets 5–7, usually ± overlapping, largest mostly oblanceolate to narrowly obovate, 2–6 (–9) × 1.5–2.5 cm, margins flat to ± revolute, distal 3/4 to nearly whole length evenly incised 3/4+ to midvein, undivided medial blade 2–4 mm wide, teeth (5–) 6–10 per side, not secondarily toothed, linear, 3–13 (–20) mm, surfaces usually ± dissimilar, abaxial grayish to whitish, long hairs abundant to dense, crisped hairs sparse to abundant, cottony hairs usually absent, glands absent or sparse, adaxial green to grayish, long hairs ± abundant, rarely dense, crisped hairs absent or sparse, cottony hairs absent, glands absent or sparse. Flowers: epicalyx bractlets 2.5–5 × 0.5–1.2 (–1.5) mm; hypanthium 3.5–5 mm diam.; petals 4–7 (–8) × (3–) 5–10 mm; carpels (20–) 30–50, styles 1.5–3 mm. Achenes 1–1.3 mm. 2n = 42.


Phenology: Flowering summer.
Habitat: Vernally moist meadows, often semialkaline, dry gravelly flats and slopes, in sagebrush communities, aspen and conifer woodlands
Elevation: 1300–3300 m

Distribution

V9 227-distribution-map.jpg

Calif., Colo., Idaho, Mont., Nev., Oreg., Utah, Wyo.

Discussion

Variety elmeri is the characteristic representative of Potentilla gracilis in the Great Basin, distinguished by deeply and evenly pectinate-toothed leaflets with a very narrow undivided medial blade. Leaflet surfaces are only somewhat dissimilar, most often grayish with abundant crisped (versus cottony) hairs abaxially. Basal leaves tend to be shorter than in other varieties, and petioles are most often tightly strigose. Intergradation with other varieties is common; populations beyond the Great Basin are problematic.

Selected References

None.

Lower Taxa

None.

"dm" is not declared as a valid unit of measurement for this property.

stiff +  and weak +
grayish +  and whitish +
1mm +  and 1.3mm +
green +  and grayish +
stiff +  and weak +
0.12 cm1.2 mm <br />0.0012 m <br /> (0.16 cm1.6 mm <br />0.0016 m <br />) +
0.06 cm0.6 mm <br />6.0e-4 m <br /> (0.12 cm1.2 mm <br />0.0012 m <br />) +
Barbara Ertter +  and James L. Reveal +
(Rydberg) Jepson +
separate +  and overlapping +
9 cm90 mm <br />0.09 m <br /> (11 cm110 mm <br />0.11 m <br />) +
0.7 cm7 mm <br />0.007 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
2 cm20 mm <br />0.02 m <br /> (9 cm90 mm <br />0.09 m <br />) +
1.5 cm15 mm <br />0.015 m <br /> (4 cm40 mm <br />0.04 m <br />) +
subpalmate +  and palmate +
40 cm400 mm <br />0.4 m <br /> (55 cm550 mm <br />0.55 m <br />) +
4 cm40 mm <br />0.04 m <br /> (40 cm400 mm <br />0.4 m <br />) +
Potentilla elmeri +
25 cm250 mm <br />0.25 m <br /> (30 cm300 mm <br />0.3 m <br />) +
less cordate or reniform +  and narrowly elliptic +
0.5 cm5 mm <br />0.005 m <br /> (25 cm250 mm <br />0.25 m <br />) +
coriaceous +
reduced +
free +  and distinct +
hairy +  and glabrous +
Elmer’s or combleaf cinquefoil +
Calif. +, Colo. +, Idaho +, Mont. +, Nev. +, Oreg. +, Utah +  and Wyo. +
not +  and aggregated +
1300–3300 m +
0.6 cm6 mm <br />0.006 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
0.12 cm1.2 mm <br />0.0012 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
linear +  and narrowly lanceolate or elliptic +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.12 cm1.2 mm <br />0.0012 m <br />) +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
0.8 cm8 mm <br />0.008 m <br /> (2 cm20 mm <br />0.02 m <br />) +
unisexual +  and bisexual +
not +  and aggregated +
deciduous +
on style scar +, hairy +  and glabrous +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.26 cm2.6 mm <br />0.0026 m <br />) +
inconspicuous +  and obscured +
Vernally moist meadows, often semialkaline, dry gravelly flats and slopes, in sagebrush communities, aspen and conifer woodlands +
stoloniferous +  and short-rhizomatous +
0.2 cm2 mm <br />0.002 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
blue-green +, green +  and grayish +
straight +
ascending +  and spreading +
sparse +  and abundant +
crisped +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
0.7 cm7 mm <br />0.007 m <br /> (1 cm10 mm <br />0.01 m <br />) +
0.35 cm3.5 mm <br />0.0035 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
(4-)10-50(-60)-flowered +
crassinucellate +
bipinnate +, odd-pinnate +, subpalmate +, palmate +  and ternate +
opposite +  and alternate +
stiff +  and weak +
distal +  and up-to-distal +
marcescent +
well-developed +
persistent +
cauline +  and basal +
reduced +
6 cm60 mm <br />0.06 m <br /> (9 cm90 mm <br />0.09 m <br />) +
2 cm20 mm <br />0.02 m <br /> (6 cm60 mm <br />0.06 m <br />) +
mostly oblanceolate +  and narrowly obovate +
1.5 cm15 mm <br />0.015 m <br /> (?) +  and 2.5 cm25 mm <br />0.025 m <br /> (?) +
linear +  and oblanceolate +
palmate +  and pinnate +
blue-green +, green +  and grayish +
stiff +  and weak +
incised +, flat +  and more or less revolute +
0.2 cm2 mm <br />0.002 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
blue-green +, green +  and grayish +
stiff +  and weak +
inferior +  and superior +
clustered +, biseriate +  and superposed +
3 cm30 mm <br />0.03 m <br /> (6.5 cm65 mm <br />0.065 m <br />) +
0.3 cm3 mm <br />0.003 m <br /> (3 cm30 mm <br />0.03 m <br />) +
0.7 cm7 mm <br />0.007 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
1 cm10 mm <br />0.01 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
0.3 cm3 mm <br />0.003 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
free +  and distinct +
0.4 cm4 mm <br />0.004 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
usually longer +
0.4 cm4 mm <br />0.004 m <br /> (1 cm10 mm <br />0.01 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (1 cm10 mm <br />0.01 m <br />) +
15 cm150 mm <br />0.15 m <br /> (25 cm250 mm <br />0.25 m <br />) +
2 cm20 mm <br />0.02 m <br /> (15 cm150 mm <br />0.15 m <br />) +
Flowering summer. +
adnate +, free +, connate +  and distinct +
subpinnate +, palmate +  and subpalmate +
not flagelliform +
2 cm20 mm <br />0.02 m <br /> (40 cm400 mm <br />0.4 m <br />) +
not arillate +
0.8 cm8 mm <br />0.008 m <br /> (1 cm10 mm <br />0.01 m <br />) +
persistent +
free +  and distinct +
erect +  and spreading +
lanceolate +  and broadly ovate or deltate +
0.4 cm4 mm <br />0.004 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
stiff +  and weak +
Endemic +  and Illustrated +
free +  and distinct +
not rooting +
reddish +  and green +
(1.5-)2-4 times basal leaves +
ascending +  and erect +
1 +  and many +
not flagelliform +
palmate +  and pinnate +
linear;ovate +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
deciduous +
distinct +
subapical +
papillate-swollen +  and tapered +
rough-thickened +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
blue-green +, green +  and grayish +
stiff +  and weak +
incised +, flat +  and more or less revolute +
P. bakeri +, P. flabelliformis var. inyoensis +, P. pectinisecta +  and P. pectinisecta var. comosa +
not fleshy-thickened +
Potentilla gracilis var. elmeri +
Potentilla gracilis +
variety +
1.3 cm13 mm <br />0.013 m <br /> (2 cm20 mm <br />0.02 m <br />) +
blue-green +, green +  and grayish +
stiff +  and weak +
6 +  and 10 +
toothed +
0.3 cm3 mm <br />0.003 m <br /> (1.3 cm13 mm <br />0.013 m <br />) +
inconspicuous +
enlarged +
biennial +, annual +  and perennial +
herb +  and matted +