Potentilla gracilis

Douglas ex Hooker

Bot. Mag. 57: plate 2984. 1830.

Common names: Slender cinquefoil potentille grêle
EndemicIllustrated
Treatment appears in FNA Volume 9. Treatment on page 153. Mentioned on page 143, 150, 151, 152, 154, 155, 156, 157, 158.

Glands usually absent or inconspicuous, uncolored. Stems ascending to erect, (0.5–) 2–7 (–12) dm. Basal leaves usually not in ranks, palmate, rarely subpalmate, 4–40 (–55) cm; petiole (2–) 3–25 (–45) cm, long hairs sparse to dense, appressed to spreading, 0.5–2 (–3) mm, weak to stiff, short hairs absent, crisped hairs usually absent, cottony hairs absent or sparse to abundant (var. owyheensis), glands absent or sparse; leaflets 5–9, on tip, rarely less than distal 1/10, of leaf axis, separate to overlapping, largest ones oblanceolate to elliptic or obovate, (1.5–) 2–9 (–11) × (0.7–) 1.5–4 cm, margins flat to revolute, distal (2/3–) 3/4 to nearly whole length evenly to unevenly incised 1/4–3/4+ to midvein, undivided medial blade 2–20 mm wide, teeth (4–) 6–10 (–11) per side (sometimes secondarily toothed), linear to broadly lanceolate, 2–20 (–35) mm, surfaces ± similar to strongly dissimilar, abaxial green or pale green to white, usually not glaucous, long hairs sparse to abundant (sometimes mostly on veins), short-crisped hairs absent or sparse to abundant, cottony hairs absent or sparse to dense, glands usually absent or obscured to sparse, sometimes common, adaxial dark green to grayish, long hairs sparse to ± abundant, rarely absent or dense, short-crisped hairs absent or sparse, cottony hairs absent or sparse to abundant (var. owyheensis), glands usually absent or sparse to sometimes common. Cauline leaves 1–4 (–5). Inflorescences (4–) 10–50 (–60) -flowered. Pedicels 0.3–3 cm. Flowers: epicalyx bractlets linear to narrowly lanceolate or elliptic, 2–6 (–8) × 0.5–1.5 (–2.5) mm, hairs sparse to abundant, rarely dense, ± appressed to ascending, rarely spreading, glands usually absent or inconspicuous; hypanthium 3.5–6 mm diam.; sepals 4–8 (–10) mm, apex acute to long acuminate; petals (3–) 4–10 (–11) × (3–) 4–10 (–12) mm; filaments (1–) 1.5–2.5 (–3.5) mm, anthers 0.6–1.2 (–1.6) mm; carpels (15–) 20–50, styles ± tapered, papillate-swollen proximally, (1–) 1.5–2.5 (–3) mm. Achenes 1–1.8 mm.

Distribution

V9 224-distribution-map.jpg

Alta., B.C., Man., N.W.T., Ont., P.E.I., Que., Sask., Yukon, Alaska, Ariz., Calif., Colo., Idaho, Mich., Minn., Mont., N.Dak., Nev., Oreg., S.Dak., Utah, Wash., Wis., Wyo.

Discussion

Varieties 5 (5 in the flora).

Because variation within Potentilla gracilis and other members of sect. Graciles is still far from resolved, taxonomic recognition is given to only the most distinctive extremes and that mostly at the varietal level. The goal is to call attention to areas that need study rather than to provide a full resolution, and even this leaves out some large, widespread populations in both keys and descriptions. The taxonomic placement of some available names, such as P. glomerata A. Nelson, P. longipedunculata Rydberg, and P. macropetala Rydberg, likewise remains unresolved.

The vouchers for the report by M. L. Fernald (1950) of Potentilla gracilis and P. pulcherrima from New Hampshire are presumably specimens collected in Coos County by A. S. Pease (NEBC). The three collections differ from one another, corresponding most closely to var. fastigiata, var. flabelliformis, and P. pulcherrima. The presence of such diversity in the same meadow, all as disjunct occurrences, suggests an artificial, non-persisting introduction.

Selected References

None.

Key

1 Petioles: cottony hairs usually abundant, rarely sparse. Potentilla gracilis var. owyheensis
1 Petioles: cottony hairs usually absent > 2
2 Leaflets incised 1/4–1/2 to midvein, undivided medial blade 5–20 mm wide, teeth narrowly lanceolate to ovate > 3
2 Leaflets incised 3/4+ to midvein, undivided medial blade 2–6 mm wide, teeth linear > 4
3 Leaflet surfaces usually strongly dissimilar, abaxial white, sometimes grayish, crisped-cottony hairs abundant to dense, rarely sparse, adaxial green; near coastal n California to Alaska. Potentilla gracilis var. gracilis
3 Leaflet surfaces ± similar to ± dissimilar, abaxial pale green to greenish gray, crisped hairs absent or sparse to abundant, cottony hairs usually absent, adaxial green to grayish green; widespread. Potentilla gracilis var. fastigiata
4 Leaflet surfaces usually ± dissimilar, abaxial grayish to whitish, long hairs abundant to dense, crisped hairs sparse to abundant, cottony hairs usually absent, adaxial green to grayish green, teeth 3–13(–20) mm, not secondarily toothed; petioles 2–15(–25) cm. Potentilla gracilis var. elmeri
4 Leaflet surfaces strongly dissimilar, abaxial white, long hairs sparse to common (mostly on veins), cottony or crisped-cottony hairs dense, straight hairs dense, mostly on veins, adaxial usually dark green, sometimes grayish green, teeth (8–)10–20(–35) mm, often secondarily toothed; petioles (7–)10–25(–30) cm. Potentilla gracilis var. flabelliformis

"dm" is not declared as a valid unit of measurement for this property.

... more about "Potentilla gracilis"
stiff +  and weak +
1mm +  and 1.8mm +
blue-green +, green +  and grayish +
stiff +  and weak +
0.12 cm1.2 mm <br />0.0012 m <br /> (0.16 cm1.6 mm <br />0.0016 m <br />) +
0.06 cm0.6 mm <br />6.0e-4 m <br /> (0.12 cm1.2 mm <br />0.0012 m <br />) +
Barbara Ertter +  and James L. Reveal +
Douglas ex Hooker +
separate +  and overlapping +
9 cm90 mm <br />0.09 m <br /> (11 cm110 mm <br />0.11 m <br />) +
0.7 cm7 mm <br />0.007 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
2 cm20 mm <br />0.02 m <br /> (9 cm90 mm <br />0.09 m <br />) +
1.5 cm15 mm <br />0.015 m <br /> (4 cm40 mm <br />0.04 m <br />) +
subpalmate +  and palmate +
40 cm400 mm <br />0.4 m <br /> (55 cm550 mm <br />0.55 m <br />) +
4 cm40 mm <br />0.04 m <br /> (40 cm400 mm <br />0.4 m <br />) +
Graciles +
25 cm250 mm <br />0.25 m <br /> (30 cm300 mm <br />0.3 m <br />) +
less cordate or reniform +  and narrowly elliptic +
0.5 cm5 mm <br />0.005 m <br /> (25 cm250 mm <br />0.25 m <br />) +
coriaceous +
reduced +
free +  and distinct +
hairy +  and glabrous +
Slender cinquefoil +  and potentille grêle +
Alta. +, B.C. +, Man. +, N.W.T. +, Ont. +, P.E.I. +, Que. +, Sask. +, Yukon +, Alaska +, Ariz. +, Calif. +, Colo. +, Idaho +, Mich. +, Minn. +, Mont. +, N.Dak. +, Nev. +, Oreg. +, S.Dak. +, Utah +, Wash. +, Wis. +  and Wyo. +
not +  and aggregated +
0.6 cm6 mm <br />0.006 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
linear +  and narrowly lanceolate or elliptic +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
0.8 cm8 mm <br />0.008 m <br /> (2 cm20 mm <br />0.02 m <br />) +
unisexual +  and bisexual +
not +  and aggregated +
deciduous +
on style scar +, hairy +  and glabrous +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.26 cm2.6 mm <br />0.0026 m <br />) +
inconspicuous +  and obscured +
stoloniferous +  and short-rhizomatous +
0.2 cm2 mm <br />0.002 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
blue-green +, green +  and grayish +
straight +
dense +  and sparse +
weak +  and stiff +
spreading +, less appressed +  and ascending +
sparse +  and abundant +
short-crisped +  and crisped +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
0.7 cm7 mm <br />0.007 m <br /> (1 cm10 mm <br />0.01 m <br />) +
0.35 cm3.5 mm <br />0.0035 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
(4-)10-50(-60)-flowered +
crassinucellate +
bipinnate +, odd-pinnate +, subpalmate +, palmate +  and ternate +
opposite +  and alternate +
stiff +  and weak +
distal +  and up-to-distal +
marcescent +
well-developed +
persistent +
cauline +  and basal +
reduced +
narrowly oblanceolate +  and obovate elliptic oblong cuneate or flabellate +
linear +  and oblanceolate +
palmate +  and pinnate +
blue-green +, green +  and grayish +
stiff +  and weak +
flat +  and revolute +
0.2 cm2 mm <br />0.002 m <br /> (2 cm20 mm <br />0.02 m <br />) +
blue-green +, green +  and grayish +
stiff +  and weak +
inferior +  and superior +
clustered +, biseriate +  and superposed +
3 cm30 mm <br />0.03 m <br /> (6.5 cm65 mm <br />0.065 m <br />) +
0.3 cm3 mm <br />0.003 m <br /> (3 cm30 mm <br />0.03 m <br />) +
1 cm10 mm <br />0.01 m <br /> (1.1 cm11 mm <br />0.011 m <br />) +
1 cm10 mm <br />0.01 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
1 cm10 mm <br />0.01 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
free +  and distinct +
0.4 cm4 mm <br />0.004 m <br /> (1 cm10 mm <br />0.01 m <br />) +
usually longer +
0.4 cm4 mm <br />0.004 m <br /> (1 cm10 mm <br />0.01 m <br />) +
0.4 cm4 mm <br />0.004 m <br /> (1 cm10 mm <br />0.01 m <br />) +
25 cm250 mm <br />0.25 m <br /> (45 cm450 mm <br />0.45 m <br />) +
3 cm30 mm <br />0.03 m <br /> (25 cm250 mm <br />0.25 m <br />) +
adnate +, free +, connate +  and distinct +
subpinnate +, palmate +  and subpalmate +
not flagelliform +
2 cm20 mm <br />0.02 m <br /> (40 cm400 mm <br />0.4 m <br />) +
not arillate +
0.8 cm8 mm <br />0.008 m <br /> (1 cm10 mm <br />0.01 m <br />) +
persistent +
free +  and distinct +
erect +  and spreading +
lanceolate +  and broadly ovate or deltate +
0.4 cm4 mm <br />0.004 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
stiff +  and weak +
Endemic +  and Illustrated +
free +  and distinct +
not rooting +
reddish +  and green +
(1.5-)2-4 times basal leaves +
ascending +  and erect +
1 +  and many +
not flagelliform +
palmate +  and pinnate +
linear;ovate +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
deciduous +
distinct +
subapical +
papillate-swollen +  and tapered +
rough-thickened +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
blue-green +, green +  and grayish +
stiff +  and weak +
incised +, flat +  and more or less revolute +
Potentilla sect. Candidae +, Potentilla sect. Heptaphyllae +, Potentilla sect. Longipedunculatae +, Potentilla sect. Nuttallianae +, Potentilla sect. Pectinisectae +  and Potentilla sect. Permolles +
not fleshy-thickened +
Potentilla gracilis +
Potentilla sect. Graciles +
species +
2 cm20 mm <br />0.02 m <br /> (3.5 cm35 mm <br />0.035 m <br />) +
blue-green +, green +  and grayish +
stiff +  and weak +
6 +  and 10 +
linear +  and broadly lanceolate +
0.2 cm2 mm <br />0.002 m <br /> (2 cm20 mm <br />0.02 m <br />) +
inconspicuous +
enlarged +
biennial +, annual +  and perennial +
herb +  and matted +