Potentilla gracilis var. fastigiata

(Nuttall) S. Watson

Proc. Amer. Acad. Arts 8: 557. 1873.

Common names: Nuttall’s or Hall’s cinquefoil potentille de Nuttall
Endemic
Basionyms: Potentilla fastigiata Nuttall in J. Torrey and A. Gray, Fl. N. Amer. 1: 440. 1840
Synonyms: P. angustata Rydberg P. blaschkeana var. permollis (Rydberg) Th. Wolf P. etomentosa Rydberg P. etomentosa subsp. hallii (Rydberg) Abrams P. fastigiata var. jucunda (A. Nelson) Soják P. fastigiata var. permollis (Rydberg) Soják P. gracilis var. glabrata (Lehmann) C. L. Hitchcock P. gracilis var. hallii (Rydberg) Th. Wolf P. gracilis subsp. nuttallii (Lehmann) D. D. Keck P. gracilis var. nuttallii (Lehmann) E. Sheldon P. gracilis var. permollis (Rydberg) C. L. Hitchcock P. gracilis var. rigida S. Watson P. indiges P. jucunda P. nuttallii P. nuttallii var. glabrata Lehmann
Treatment appears in FNA Volume 9. Treatment on page 155. Mentioned on page 154.

Stems (0.5–) 1.5–7 (–9) dm. Basal leaves: petiole (2–) 3–20 (–45) cm, long hairs common to dense, appressed to spreading, 0.5–2 (–3) mm, stiff, crisped hairs usually absent, sometimes sparse to common, cottony hairs absent, glands absent or sparse; leaflets 5–7 (–9), separate to ± overlapping, largest oblanceolate to elliptic or narrowly obovate, (1.5–) 2–7 (–10) × (0.7–) 1.5–3.5 cm, margins flat to slightly revolute, distal (2/3–) 3/4 to nearly whole length ± evenly incised 1/4–1/2 to midvein, undivided medial blade 6–20 mm wide, teeth 5–9 (–11) per side, usually not secondarily toothed, narrowly lanceolate to ovate, 2–10 mm, surfaces ± similar to ± dissimilar, abaxial pale green to greenish gray, long hairs sparse to abundant (mostly on veins), crisped hairs absent or sparse to abundant, cottony hairs usually absent, glands absent or sparse, rarely common, adaxial green to grayish green, long hairs sparse to common, sometimes absent, crisped and cottony hairs absent, glands absent or sparse, rarely common. Flowers: epicalyx bractlets 2–6.5 (–7) × 0.5–1.5 (–2.5) mm; hypanthium 4–6 mm diam.; petals (3–) 4–8 (–10) × (3–) 4–10 (–12) mm; carpels (15–) 30–50, styles 1.5–2.5 (–3) mm. Achenes 1.4–1.8 mm.


Phenology: Flowering summer.
Habitat: Vernally moist to dry meadows or gravelly flats and slopes, in grassland and sagebrush communities, oak, aspen, or conifer woodlands
Elevation: (10–)100–3500 m

Distribution

V9 226-distribution-map.jpg

Alta., B.C., Man., N.W.T., Ont., P.E.I., Que., Sask., Yukon, Alaska, Ariz., Calif., Colo., Idaho, Minn., Mont., Nev., N.Dak., Oreg., S.Dak., Utah, Wash., Wyo.

Discussion

Variety fastigiata was commonly known as var. nuttallii until N. H. Holmgren (1997b) noted that var. fastigiata had priority at varietal rank, due to the publication of Potentilla holopetala Turczaninow var. fastigiata (Nuttall) Lehmann. As used here, the variety encompasses most collections of P. gracilis with moderately and evenly toothed leaflets that have at most somewhat dissimilar leaflet surfaces that usually lack cottony hairs. As such, the variety is essentially the catch-all category for most variation within P. gracilis that is not assigned elsewhere.

Within this broad circumscription are at least three candidates for taxonomic recognition. First is var. fastigiata in the narrow sense, consisting of relatively large, hairy plants found throughout the range given above. The hallii phase, found mostly in the mountains of California, is shorter on average, and leaflets tend to be broader, less hairy, and less deeply incised. Glabrate populations that are common from eastern Oregon to the northern Rocky Mountains, some possibly indicating introgression from Potentilla glaucophylla, represent the nuttallii or glabrata phase.

Occurrences in Quebec and Prince Edward Island, and at least some other occurrences in eastern provinces and states, are most likely introduced. Eastern populations of var. fastigiata in general tend to be relatively hairy and to intergrade with Potentilla pulcherrima.

Selected References

None.

Lower Taxa

None.

"dm" is not declared as a valid unit of measurement for this property.

stiff +  and weak +
pale green +  and greenish gray +
1.4mm +  and 1.8mm +
green +  and grayish green +
blue-green +, green +  and grayish +
stiff +  and weak +
0.12 cm1.2 mm <br />0.0012 m <br /> (0.16 cm1.6 mm <br />0.0016 m <br />) +
0.06 cm0.6 mm <br />6.0e-4 m <br /> (0.12 cm1.2 mm <br />0.0012 m <br />) +
Barbara Ertter +  and James L. Reveal +
(Nuttall) S. Watson +
separate +  and overlapping +
9 cm90 mm <br />0.09 m <br /> (11 cm110 mm <br />0.11 m <br />) +
0.7 cm7 mm <br />0.007 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
2 cm20 mm <br />0.02 m <br /> (9 cm90 mm <br />0.09 m <br />) +
1.5 cm15 mm <br />0.015 m <br /> (4 cm40 mm <br />0.04 m <br />) +
subpalmate +  and palmate +
40 cm400 mm <br />0.4 m <br /> (55 cm550 mm <br />0.55 m <br />) +
4 cm40 mm <br />0.04 m <br /> (40 cm400 mm <br />0.4 m <br />) +
Potentilla fastigiata +
25 cm250 mm <br />0.25 m <br /> (30 cm300 mm <br />0.3 m <br />) +
less cordate or reniform +  and narrowly elliptic +
0.5 cm5 mm <br />0.005 m <br /> (25 cm250 mm <br />0.25 m <br />) +
coriaceous +
reduced +
free +  and distinct +
hairy +  and glabrous +
Nuttall’s or Hall’s cinquefoil +  and potentille de Nuttall +
Alta. +, B.C. +, Man. +, N.W.T. +, Ont. +, P.E.I. +, Que. +, Sask. +, Yukon +, Alaska +, Ariz. +, Calif. +, Colo. +, Idaho +, Minn. +, Mont. +, Nev. +, N.Dak. +, Oreg. +, S.Dak. +, Utah +, Wash. +  and Wyo. +
not +  and aggregated +
(10–)100–3500 m +
0.65 cm6.5 mm <br />0.0065 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (0.65 cm6.5 mm <br />0.0065 m <br />) +
linear +  and narrowly lanceolate or elliptic +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
0.8 cm8 mm <br />0.008 m <br /> (2 cm20 mm <br />0.02 m <br />) +
unisexual +  and bisexual +
not +  and aggregated +
deciduous +
on style scar +, hairy +  and glabrous +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.26 cm2.6 mm <br />0.0026 m <br />) +
inconspicuous +  and obscured +
Vernally moist to dry meadows or gravelly flats and slopes, in grassland and sagebrush communities, oak, aspen, or conifer woodlands +
stoloniferous +  and short-rhizomatous +
0.2 cm2 mm <br />0.002 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
blue-green +, green +  and grayish +
straight +
appressed +  and spreading +
sparse +  and common +
crisped +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
0.7 cm7 mm <br />0.007 m <br /> (1 cm10 mm <br />0.01 m <br />) +
0.4 cm4 mm <br />0.004 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
(4-)10-50(-60)-flowered +
crassinucellate +
bipinnate +, odd-pinnate +, subpalmate +, palmate +  and ternate +
opposite +  and alternate +
stiff +  and weak +
distal +  and up-to-distal +
marcescent +
well-developed +
persistent +
cauline +  and basal +
reduced +
separate +  and more or less overlapping +
7 cm70 mm <br />0.07 m <br /> (10 cm100 mm <br />0.1 m <br />) +
0.7 cm7 mm <br />0.007 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
2 cm20 mm <br />0.02 m <br /> (7 cm70 mm <br />0.07 m <br />) +
oblanceolate +  and elliptic or narrowly obovate +
1.5 cm15 mm <br />0.015 m <br /> (?) +  and 3.5 cm35 mm <br />0.035 m <br /> (?) +
linear +  and oblanceolate +
palmate +  and pinnate +
blue-green +, green +  and grayish +
stiff +  and weak +
flat +  and slightly revolute +
0.6 cm6 mm <br />0.006 m <br /> (2 cm20 mm <br />0.02 m <br />) +
blue-green +, green +  and grayish +
stiff +  and weak +
inferior +  and superior +
clustered +, biseriate +  and superposed +
3 cm30 mm <br />0.03 m <br /> (6.5 cm65 mm <br />0.065 m <br />) +
0.3 cm3 mm <br />0.003 m <br /> (3 cm30 mm <br />0.03 m <br />) +
0.8 cm8 mm <br />0.008 m <br /> (1 cm10 mm <br />0.01 m <br />) +
1 cm10 mm <br />0.01 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
1 cm10 mm <br />0.01 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
free +  and distinct +
0.4 cm4 mm <br />0.004 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
usually longer +
0.4 cm4 mm <br />0.004 m <br /> (1 cm10 mm <br />0.01 m <br />) +
0.4 cm4 mm <br />0.004 m <br /> (1 cm10 mm <br />0.01 m <br />) +
20 cm200 mm <br />0.2 m <br /> (45 cm450 mm <br />0.45 m <br />) +
3 cm30 mm <br />0.03 m <br /> (20 cm200 mm <br />0.2 m <br />) +
Flowering summer. +
adnate +, free +, connate +  and distinct +
subpinnate +, palmate +  and subpalmate +
not flagelliform +
2 cm20 mm <br />0.02 m <br /> (40 cm400 mm <br />0.4 m <br />) +
Proc. Amer. Acad. Arts +
not arillate +
0.8 cm8 mm <br />0.008 m <br /> (1 cm10 mm <br />0.01 m <br />) +
persistent +
free +  and distinct +
erect +  and spreading +
lanceolate +  and broadly ovate or deltate +
0.4 cm4 mm <br />0.004 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
stiff +  and weak +
free +  and distinct +
not rooting +
reddish +  and green +
(1.5-)2-4 times basal leaves +
ascending +  and erect +
1 +  and many +
not flagelliform +
palmate +  and pinnate +
linear;ovate +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
deciduous +
distinct +
subapical +
papillate-swollen +  and tapered +
rough-thickened +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
blue-green +, green +  and grayish +
stiff +  and weak +
incised +, flat +  and more or less revolute +
P. angustata +, P. blaschkeana var. permollis +, P. etomentosa +, P. etomentosa subsp. hallii +, P. fastigiata var. jucunda +, P. fastigiata var. permollis +, P. gracilis var. glabrata +, P. gracilis var. hallii +, P. gracilis subsp. nuttallii +, P. gracilis var. nuttallii +, P. gracilis var. permollis +, P. gracilis var. rigida +, P. indiges +, P. jucunda +, P. nuttallii +  and P. nuttallii var. glabrata +
not fleshy-thickened +
Potentilla gracilis var. fastigiata +
Potentilla gracilis +
variety +
2 cm20 mm <br />0.02 m <br /> (3.5 cm35 mm <br />0.035 m <br />) +
blue-green +, green +  and grayish +
stiff +  and weak +
5 +  and 9 +
narrowly lanceolate +  and ovate +
0.2 cm2 mm <br />0.002 m <br /> (1 cm10 mm <br />0.01 m <br />) +
inconspicuous +
enlarged +
biennial +, annual +  and perennial +
herb +  and matted +