Potentilla gracilis var. gracilis

Endemic
Synonyms: Potentilla alaskana Rydberg P. blaschkeana Turczaninow ex Lehmann
Treatment appears in FNA Volume 9. Treatment on page 154. Mentioned on page 155, 157, 158.

Stems (2–) 4–10 (–12) dm. Basal leaves: petiole (7–) 10–25 cm, long hairs abundant, usually spreading, sometimes appressed, 0.5–2 (–3) mm, stiff, crisped and cottony hairs usually absent, glands absent; leaflets (5–) 7–9, ± separate, largest oblanceolate to narrowly obovate, 3–9 × (1.3–) 1.5–3 (–3.5) cm, margins flat to ± revolute, nearly whole length ± evenly incised 1/3–1/2 to midvein, undivided medial blade 5–12 mm wide, teeth 7–10 (–11) per side, sometimes secondarily toothed, narrowly lanceolate to ovate, 3–8 mm, surfaces usually strongly dissimilar, abaxial white, sometimes grayish, long hairs abundant (mostly on veins), crisped-cottony hairs abundant to dense, rarely sparse, glands absent or sparse, adaxial green, straight hairs sparse, rarely absent, crisped and cottony hairs absent, glands absent. Flowers: epicalyx bractlets 3–5 (–6) × 0.5–1.1 mm; hypanthium 4–6 mm diam.; petals (5–) 7–10 × 5–12 mm; carpels 20–40, styles 2–3 mm. Achenes 1.2–1.8 mm.


Phenology: Flowering summer.
Habitat: Moist meadows, usually in oak and conifer woodlands
Elevation: 100–500(–1100) m

Distribution

V9 225-distribution-map.jpg

B.C., Alaska, Calif., Oreg., Wash.

Discussion

Variety gracilis is a relatively tall plant with serrate, narrow, strongly bicolor leaves, confined mainly to meadows west of the Cascade Mountains, from northwestern California northward through western Oregon to southern Washington. From there, an expression with increasingly broad leaflets transitional to var. fastigiata, represented by Potentilla alaskana and P. blaschkeana, extends in the coastal mountains to southern Alaska.

References to var. gracilis from the Canadian prairie provinces (B. Boivin 1967–1979) correspond to the Potentilla filipes Rydberg extreme of P. pulcherrima. The two are superficially similar; var. gracilis has more sharply and deeply serrate leaflets and lacks the conspicuous glands and contrasting leaflet veins of the latter. The inclusion of Saskatchewan in the range of var. gracilis by H. J. Scoggan (1978–1979, part 3) is most likely based on var. flabelliformis or its intergrades with other varieties.

Selected References

None.

Lower Taxa

None.

"dm" is not declared as a valid unit of measurement for this property.

stiff +  and weak +
grayish +  and white +
1.2mm +  and 1.8mm +
stiff +  and weak +
0.12 cm1.2 mm <br />0.0012 m <br /> (0.16 cm1.6 mm <br />0.0016 m <br />) +
0.06 cm0.6 mm <br />6.0e-4 m <br /> (0.12 cm1.2 mm <br />0.0012 m <br />) +
Barbara Ertter +  and James L. Reveal +
Douglas ex Hooker +
separate +  and overlapping +
9 cm90 mm <br />0.09 m <br /> (11 cm110 mm <br />0.11 m <br />) +
0.7 cm7 mm <br />0.007 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
2 cm20 mm <br />0.02 m <br /> (9 cm90 mm <br />0.09 m <br />) +
1.5 cm15 mm <br />0.015 m <br /> (4 cm40 mm <br />0.04 m <br />) +
subpalmate +  and palmate +
40 cm400 mm <br />0.4 m <br /> (55 cm550 mm <br />0.55 m <br />) +
4 cm40 mm <br />0.04 m <br /> (40 cm400 mm <br />0.4 m <br />) +
Graciles +
25 cm250 mm <br />0.25 m <br /> (30 cm300 mm <br />0.3 m <br />) +
less cordate or reniform +  and narrowly elliptic +
0.5 cm5 mm <br />0.005 m <br /> (25 cm250 mm <br />0.25 m <br />) +
coriaceous +
reduced +
free +  and distinct +
hairy +  and glabrous +
B.C. +, Alaska +, Calif. +, Oreg. +  and Wash. +
not +  and aggregated +
100–500(–1100) m +
0.5 cm5 mm <br />0.005 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
0.3 cm3 mm <br />0.003 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
linear +  and narrowly lanceolate or elliptic +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.11 cm1.1 mm <br />0.0011 m <br />) +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
0.8 cm8 mm <br />0.008 m <br /> (2 cm20 mm <br />0.02 m <br />) +
unisexual +  and bisexual +
not +  and aggregated +
deciduous +
on style scar +, hairy +  and glabrous +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.26 cm2.6 mm <br />0.0026 m <br />) +
inconspicuous +  and obscured +
Moist meadows, usually in oak and conifer woodlands +
stoloniferous +  and short-rhizomatous +
0.2 cm2 mm <br />0.002 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
blue-green +, green +  and grayish +
straight +
cottony +  and crisped-cottony +
crisped +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
0.7 cm7 mm <br />0.007 m <br /> (1 cm10 mm <br />0.01 m <br />) +
0.4 cm4 mm <br />0.004 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
(4-)10-50(-60)-flowered +
crassinucellate +
bipinnate +, odd-pinnate +, subpalmate +, palmate +  and ternate +
opposite +  and alternate +
stiff +  and weak +
distal +  and up-to-distal +
marcescent +
well-developed +
persistent +
cauline +  and basal +
reduced +
3 cm30 mm <br />0.03 m <br /> (3.5 cm35 mm <br />0.035 m <br />) +
3 cm30 mm <br />0.03 m <br /> (9 cm90 mm <br />0.09 m <br />) +
oblanceolate +  and narrowly obovate +
1.5 cm15 mm <br />0.015 m <br /> (?) +  and 3 cm30 mm <br />0.03 m <br /> (?) +
linear +  and oblanceolate +
palmate +  and pinnate +
blue-green +, green +  and grayish +
stiff +  and weak +
1/3 +  and 1/2 +
incised +, flat +  and more or less revolute +
0.5 cm5 mm <br />0.005 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
blue-green +, green +  and grayish +
stiff +  and weak +
inferior +  and superior +
clustered +, biseriate +  and superposed +
3 cm30 mm <br />0.03 m <br /> (6.5 cm65 mm <br />0.065 m <br />) +
0.3 cm3 mm <br />0.003 m <br /> (3 cm30 mm <br />0.03 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
1 cm10 mm <br />0.01 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
1 cm10 mm <br />0.01 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
free +  and distinct +
0.7 cm7 mm <br />0.007 m <br /> (1 cm10 mm <br />0.01 m <br />) +
usually longer +
0.4 cm4 mm <br />0.004 m <br /> (1 cm10 mm <br />0.01 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
7 cm70 mm <br />0.07 m <br /> (10 cm100 mm <br />0.1 m <br />) +
10 cm100 mm <br />0.1 m <br /> (25 cm250 mm <br />0.25 m <br />) +
Flowering summer. +
adnate +, free +, connate +  and distinct +
subpinnate +, palmate +  and subpalmate +
not flagelliform +
2 cm20 mm <br />0.02 m <br /> (40 cm400 mm <br />0.4 m <br />) +
not arillate +
0.8 cm8 mm <br />0.008 m <br /> (1 cm10 mm <br />0.01 m <br />) +
persistent +
free +  and distinct +
erect +  and spreading +
lanceolate +  and broadly ovate or deltate +
0.4 cm4 mm <br />0.004 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
stiff +  and weak +
free +  and distinct +
not rooting +
reddish +  and green +
(1.5-)2-4 times basal leaves +
ascending +  and erect +
1 +  and many +
not flagelliform +
palmate +  and pinnate +
linear;ovate +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
deciduous +
distinct +
subapical +
papillate-swollen +  and tapered +
rough-thickened +
0.2 cm2 mm <br />0.002 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
blue-green +, green +  and grayish +
stiff +  and weak +
incised +, flat +  and more or less revolute +
Potentilla alaskana +  and P. blaschkeana +
not fleshy-thickened +
Potentilla gracilis var. gracilis +
Potentilla gracilis +
variety +
2 cm20 mm <br />0.02 m <br /> (3.5 cm35 mm <br />0.035 m <br />) +
blue-green +, green +  and grayish +
stiff +  and weak +
7 +  and 10 +
narrowly lanceolate +  and ovate +
0.3 cm3 mm <br />0.003 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
inconspicuous +
enlarged +
biennial +, annual +  and perennial +
herb +  and matted +