Potentilla gracilis var. owyheensis

Ertter & Mansfield

Madroño 54: 181, fig. 1. 2007.

Common names: Owyhee cinquefoil
Endemic
Treatment appears in FNA Volume 9. Treatment on page 156. Mentioned on page 153, 154, 206.

Stems (1.5–) 3–6 dm. Basal leaves: petiole (3–) 5–15 (–19) cm, long hairs weak, sparse to abundant, loosely appressed to ascending, 1–2 mm, crisped and/or cottony hairs usually abundant, rarely sparse, glands absent or obscured; leaflets 5–7, ± overlapping, largest oblanceolate to obovate, (2–) 3.5–7 × (1–) 2–4 cm, margins flat, distal 3/4 to nearly whole length evenly to unevenly incised (1/2–) 3/4+ to midvein, undivided medial blade 4–10 mm wide, teeth 4–8 per side, often secondarily toothed, linear to lanceolate, 7–17 mm, surfaces ± similar to ± dissimilar, abaxial grayish white, long hairs sparse to common, crisped and/or cottony hairs abundant to dense, rarely sparse, glands absent or obscured, adaxial greenish gray, long hairs sparse to common, crisped and/or cottony hairs sparse to abundant, glands absent or obscured to sparse. Flowers: epicalyx bractlets 2–5 (–6) × 0.5–1.5 mm; hypanthium 3.5–5 mm diam.; petals (3–) 4–6 (–11) × 4–12 mm; carpels 25–40, styles 1.5–2.5 mm. Achenes 1.4–1.5 mm.


Phenology: Flowering summer.
Habitat: Vernally moist meadows, in sagebrush communities, aspen and conifer woodlands
Elevation: 1200–2700 m

Discussion

Variety owyheensis ranges from the Owyhee uplands of Owyhee County, Idaho, to the Ochoco Mountains of central and eastern Oregon. It is readily distinguished from other varieties in the abundance of loose cottony hairs (grading to long soft hairs) on petioles and both leaflet surfaces. The grayish white leaflets tend to be unevenly incised and secondarily toothed. Where sharing meadows with other varieties, var. owyheensis generally grows closest to the dry margins.

Comparable plants from the northern Sierra Nevada of California, represented by the type of Potentilla subvillosa Rydberg, might prove to belong here as well.

Selected References

None.

Lower Taxa

None.

"dm" is not declared as a valid unit of measurement for this property.

stiff +  and weak +
greenish gray +  and grayish white +
greenish gray +  and grayish white +
greenish gray +  and grayish white +
1.4mm +  and 1.5mm +
stiff +  and weak +
0.12 cm1.2 mm <br />0.0012 m <br /> (0.16 cm1.6 mm <br />0.0016 m <br />) +
0.06 cm0.6 mm <br />6.0e-4 m <br /> (0.12 cm1.2 mm <br />0.0012 m <br />) +
Barbara Ertter +  and James L. Reveal +
Ertter & Mansfield +
separate +  and overlapping +
9 cm90 mm <br />0.09 m <br /> (11 cm110 mm <br />0.11 m <br />) +
0.7 cm7 mm <br />0.007 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
2 cm20 mm <br />0.02 m <br /> (9 cm90 mm <br />0.09 m <br />) +
1.5 cm15 mm <br />0.015 m <br /> (4 cm40 mm <br />0.04 m <br />) +
subpalmate +  and palmate +
40 cm400 mm <br />0.4 m <br /> (55 cm550 mm <br />0.55 m <br />) +
4 cm40 mm <br />0.04 m <br /> (40 cm400 mm <br />0.4 m <br />) +
Graciles +
25 cm250 mm <br />0.25 m <br /> (30 cm300 mm <br />0.3 m <br />) +
less cordate or reniform +  and narrowly elliptic +
0.5 cm5 mm <br />0.005 m <br /> (25 cm250 mm <br />0.25 m <br />) +
coriaceous +
reduced +
free +  and distinct +
hairy +  and glabrous +
Owyhee cinquefoil +
Idaho +  and Oreg. +
not +  and aggregated +
1200–2700 m +
0.5 cm5 mm <br />0.005 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
linear +  and narrowly lanceolate or elliptic +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
0.8 cm8 mm <br />0.008 m <br /> (2 cm20 mm <br />0.02 m <br />) +
unisexual +  and bisexual +
not +  and aggregated +
deciduous +
on style scar +, hairy +  and glabrous +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.26 cm2.6 mm <br />0.0026 m <br />) +
Vernally moist meadows, in sagebrush communities, aspen and conifer woodlands +
stoloniferous +  and short-rhizomatous +
0.2 cm2 mm <br />0.002 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
blue-green +, green +  and grayish +
straight +
dense +  and sparse +
loosely appressed +  and ascending +
abundant +  and sparse +
crisped +
0.1 cm1 mm <br />0.001 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
0.7 cm7 mm <br />0.007 m <br /> (1 cm10 mm <br />0.01 m <br />) +
0.35 cm3.5 mm <br />0.0035 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
(4-)10-50(-60)-flowered +
crassinucellate +
bipinnate +, odd-pinnate +, subpalmate +, palmate +  and ternate +
opposite +  and alternate +
stiff +  and weak +
distal +  and up-to-distal +
marcescent +
well-developed +
persistent +
cauline +  and basal +
reduced +
2 cm20 mm <br />0.02 m <br /> (3.5 cm35 mm <br />0.035 m <br />) +
1 cm10 mm <br />0.01 m <br /> (2 cm20 mm <br />0.02 m <br />) +
3.5 cm35 mm <br />0.035 m <br /> (7 cm70 mm <br />0.07 m <br />) +
oblanceolate +  and obovate +
2 cm20 mm <br />0.02 m <br /> (?) +  and 4 cm40 mm <br />0.04 m <br /> (?) +
linear +  and oblanceolate +
palmate +  and pinnate +
blue-green +, green +  and grayish +
stiff +  and weak +
1/2 +  and 3/4 +
0.4 cm4 mm <br />0.004 m <br /> (1 cm10 mm <br />0.01 m <br />) +
blue-green +, green +  and grayish +
stiff +  and weak +
inferior +  and superior +
clustered +, biseriate +  and superposed +
3 cm30 mm <br />0.03 m <br /> (6.5 cm65 mm <br />0.065 m <br />) +
0.3 cm3 mm <br />0.003 m <br /> (3 cm30 mm <br />0.03 m <br />) +
0.6 cm6 mm <br />0.006 m <br /> (1.1 cm11 mm <br />0.011 m <br />) +
1 cm10 mm <br />0.01 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
1 cm10 mm <br />0.01 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
free +  and distinct +
0.4 cm4 mm <br />0.004 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
usually longer +
0.4 cm4 mm <br />0.004 m <br /> (1 cm10 mm <br />0.01 m <br />) +
0.4 cm4 mm <br />0.004 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
15 cm150 mm <br />0.15 m <br /> (19 cm190 mm <br />0.19 m <br />) +
5 cm50 mm <br />0.05 m <br /> (15 cm150 mm <br />0.15 m <br />) +
Flowering summer. +
adnate +, free +, connate +  and distinct +
subpinnate +, palmate +  and subpalmate +
not flagelliform +
2 cm20 mm <br />0.02 m <br /> (40 cm400 mm <br />0.4 m <br />) +
not arillate +
0.8 cm8 mm <br />0.008 m <br /> (1 cm10 mm <br />0.01 m <br />) +
persistent +
free +  and distinct +
erect +  and spreading +
lanceolate +  and broadly ovate or deltate +
0.4 cm4 mm <br />0.004 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
stiff +  and weak +
free +  and distinct +
not rooting +
reddish +  and green +
(1.5-)2-4 times basal leaves +
ascending +  and erect +
1 +  and many +
not flagelliform +
palmate +  and pinnate +
linear;ovate +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
deciduous +
distinct +
subapical +
papillate-swollen +  and tapered +
rough-thickened +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
blue-green +, green +  and grayish +
stiff +  and weak +
incised +, flat +  and more or less revolute +
Potentilla sect. Candidae +, Potentilla sect. Heptaphyllae +, Potentilla sect. Longipedunculatae +, Potentilla sect. Nuttallianae +, Potentilla sect. Pectinisectae +  and Potentilla sect. Permolles +
not fleshy-thickened +
Potentilla gracilis var. owyheensis +
Potentilla gracilis +
variety +
2 cm20 mm <br />0.02 m <br /> (3.5 cm35 mm <br />0.035 m <br />) +
blue-green +, green +  and grayish +
stiff +  and weak +
4 +  and 8 +
linear +  and lanceolate +
0.7 cm7 mm <br />0.007 m <br /> (1.7 cm17 mm <br />0.017 m <br />) +
inconspicuous +
enlarged +
biennial +, annual +  and perennial +
herb +  and matted +